Belongs within: Neogastropoda.
Contains: Zeatrophon, Axymene, Terefundus, Poirieria, Typhinae, Muricopsis, Favartia, Murexiella, Murexsul, Coralliophilinae, Ocenebra, Nucella, Trophon, Rapaninae, Ergalataxinae, Dermomurex, Pteropurpura, Haustellum, Pterynotus, Murex, Chicoreus, Siratus.
The Muricidae are a group of predatory marine gastropods that often possess strongly developed varices on the shell. The varices may bear prominent spines, flanges or other ornaments. Muricids possess an accessory boring organ in the sole of the foot that secretes a compound used to soften the shells of mollusc prey, allowing their radula to erode a hole through the shell. They also produce a purple secretion from the hypobranchial organ used in defense.
Within the Muricidae, members of the subfamily Muricinae are characterised by a long, straight siphonal canal and spinose or frondose varices. The Trophoninae have a similarly elongate siphonal canal but possess numerous laminate varices. The Trophoninae and the related subfamily Ocenebrinae have a similar rachidian tooth morphology but Ocenebrinae (which include forms both with and without varices) have bifid marginal teeth at maturity. Aspella is a genus of small murexes with later whorls flattened (Wilson 1994). Attiliosa species bear solid shells with a sculpture of scabrous spiral cords anteriorly and low peripheral nodules (Wilson 1994).
The royal snail
Published 2 May 2025
The carnivorous neogastropods are without question one of the most successful lineages of molluscs within the modern fauna. Among the numerous representatives of this diverse clade, perhaps the most eye-catching are to be found among the Muricidae, the murexes and oyster drills.
Over 1700 species are currently recognised among the modern muricids. They come in a wide range of sizes and shapes, some growing quite large. Many species are characterised by the development of varices, regular axial ornaments of the shell. At its most basic form, a varix is simply a thickened ridge running the length of the shell, but many murexes exhibit modifications of the varices into laminae, spines, frills, or other such extravagances. The function of such modifications (if any) remains poorly known: they are often presumed to be protective structures, though they may also provide mechanical functions such as improving weight distribution on soft sediments.
Internally, the majority of muricids are more uniform (with notable exceptions). They have a rachiglossan radula, with a single central and two lateral teeth per row. A pair of accessory salivary glands are present, as is an accessory boring organ associated with the radula. A hypobranchial gland produces a purple defensive secretion (Barco et al. 2010). Murexes are generally predators of other hard-shelled invertebrates such as barnacles or other molluscs. Acid secretions from the accessory boring organ are used to soften the shell of prey as the radula is used to scrape through to the soft flesh inside. Even if you’ve never seen a live muricid, you may well have seen their leavings: a discarded clam shell with a perfectly round hole bored into it.
Over time, the muricids have been divided between numerous subfamilies though the definitions of many have been subject to debate. Molecular studies do support a basal division between two major clades, one of which includes many of the larger and often more ornate species such as murexes and trophons. The members of the second clade are often smaller and include the species known as oyster drills. They also include the Coralliophilinae, a derived subgroup of the muricids that have become specialised parasites on corals and other anthozoans. Coralliophilines have lost the radula, salivary organs and boring organ of other muricids and instead feed suctorially. Some have gone so far as to become completely sedentary endoparasites with irregular shell forms to match.
The ways in which murexes have impinged upon human lives have also been various. A number of larger species are collected as food or for their attractive and interesting shells. Conversely, some of the oyster drills are regarded as significant pests, attacking stocks of farmed shellfish. Perhaps the most remarkable way that murexes have impacted history, however, comes from the aforementioned purple hypobranchial secretion. This is the source of the Tyrian purple dye that was so prized in Mediterranean antiquity. Because of the intense amount of time and labour that was required to obtain usable quantities of this dye, it became regarded as the preserve of wealth and power. Roman law would eventually restrict its usage to the person of the emperor, and to assume royal status become to ‘don the purple’. And all from the futile attempts at self-defense by an unfortunate snail.
Systematics of Muricidae
<==Muricidae [Muricedae, Muricoidea, Trophonidae, Typhididae]BR17
|--+--OcenebrinaeZLK11
| | |--OcenebraBR05
| | |--NucellaZLK11
| | `--Ecphora Conrad 1843 [Ecphorinae]BR05
| | |--*E. (Ecphora) quadricostata (Say 1824) [=Fusus quadricostatus]RD75
| | `--E. (Planecphora) delicata Petuch 1989P12
| `--TrophoninaeZLK11
| |--TrophonBD12
| |--Enatimene Iredale 1929W94
| | `--*E. simplex (Hedley 1903)W94 [=Trophon simplexW94, Murex simplexRD75]
| |--Gemixystus Iredale 1929W94
| | `--*G. laminatus (Petterd 1884) [=Trophon laminatus]RD75
| |--Ollaphon Iredale 1929W94
| | `--*O. molorthus (Hedley & May 1908) [=Trophon molorthus]RD75
| |--Xenotrophon Iredale 1929W94
| | `--*X. euschema (Iredale 1929) [=Trophon euschema]RD75
| |--Anatrophon Iredale 1929W94
| | |--*A. sarmentosa (Hedley & May 1908) [=Trophon sarmentosus]W94
| | `--A. latior (Verco 1909)W94
| |--Benthoxystus Iredale 1929W94
| | |--*B. columnarius (Hedley & May 1908) [=Trophon columnarius]RD75
| | `--B. recurvatus (Verco 1909)W94
| |--Enixotrophon Iredale 1929W94
| | |--*E. carduelis (Watson 1886) [=Trophon carduelis]W94
| | `--E. obtusiliratum (Schepman 1911)W94
| |--Apixystus Iredale 1929W94
| | |--*A. stimuleus (Hedley 1907) [=Trophon stimuleus]RD75
| | |--A. plicilaminatus (Verco 1909)W94
| | `--A. segmentatus (Verco 1909)W94
| |--Litozamia Iredale 1929W94
| | |--*L. rudolphi (Brazier 1894)RD75, W94 [=Peristernia rudolphiRD75]
| | |--L. brazieri (Tenison Woods 1875)W94
| | |--L. longior (Verco 1909)W94
| | `--L. petterdi (Crosse 1870)W94
| `--Boreotrophon Fischer 1884ZLK11, RD75
| |--*B. clathrata (Linné 1767)RD75 (see below for synonymy)
| |--B. bentleyiB56
| |--B. candelabrumGAS03
| |--B. triangulatusB56
| `--B. xestraZLK11
`--+--+--RapaninaeZLK11
| `--+--ErgalataxinaeZLK11
| `--Drupella Thiele 1925ZLK11, WG71
| |--D. cornus (Röding 1798)BW09 [incl. Purpura elata Blainville 1832H08, *D. elataW94]
| |--D. margariticola (Broderip in Broderip & Sowerby 1833)WBS-S15
| |--D. minuta Fujioka 1984BC01
| |--D. ochrostoma (Blainville 1832)RD75 (see below for synonymy)
| `--D. rugosa (Born 1778) [incl. D. concatenatus Lamarck 1822, D. fragum Blainville 1832]W94
`--Muricinae [Murexia]ZLK11
| i. s.: DermomurexV95
| PteropurpuraO27
| Trialatella Berry 1964PP78, RD75
| `--*T. cunninghamae Berry 1964RD75 [=Aspella (Trialatella) cunninghamaePP78]
| Attiliosa Emerson 1968W94
| |--*A. incompta (Berry 1960) [=Coralliophila incompta]RD75
| |--A. aldridgei (Usticke 1969) [=Vasum aldridgei]BC01
| |--A. carmen (Lowe 1935)PP78
| `--A. nodulifera (Sowerby 1841)W94
| HaustellumW94
| Hexaplex Swainson 1833W94, BC01 (see below for synonymy)
| |--H. cichoreum (Gmelin 1791)DM84 [=Murex cichoreumW94; incl. *H. foliacea Perry 1811W94]
| |--H. conatus (McMichael 1964) [=Murexsul conatus]W94
| |--H. fulvescens (Sowerby 1834)DM84
| |--H. kusterianus (Tapparone-Canefri 1875)DM84 [=Muricanthus kusterianusBC01]
| | |--H. k. kusterianusBC01
| | `--H. k. bozzadamii (Franchi 1990) [=Muricanthus kusterianus bozzadamii]BC01
| |--H. rileyi D’Attilio & Myers 1984BC01
| |--H. stainforthi (Reeve 1843)W94 (see below for synonymy)
| `--H. trunculus (Linné 1758)DM84
| Naquetia Jousseaume 1880W94
| |--*N. triqueter (Born 1778)W94 (see below for synonymy)
| | |--N. t. triqueterW94
| | `--N. t. vokesae Houart 1986W94
| |--N. barclayi (Reeve 1858) [incl. N. annandalei Preston 1910]W94
| |--N. cumingii (Adams 1853) (see below for synonymy)W94
| |--N. fosteri D’Attilio & Hertz 1987BC01
| `--N. permaesta (Hedley 1915) [=Pterynotus (Naquetia) permaestus]WG71
| PterynotusW94
|--MurexZLK11
`--+--+--ChicoreusZLK11
| `--Bolinus Pusch 1837S11, RD75 [=Rhinocantha Adams & Adams 1853RD75]
| |--*B. brandaris (Linnaeus 1758) [=Murex brandaris, *Rhinocantha brandaris]RD75
| `--B. cornutus (Linné 1758)LB12
`--+--SiratusZLK11
`--Muricanthus Swainson 1840ZLK11, RD75 [=Centronotus Swainson 1833 non Schneider 1801RD75]
|--*M. radix (Gmelin 1791)RD75 [=Murex radixRD75, *Centronotus radixRD75; incl. Murex melanomathosC64]
|--‘Murex’ duplex Röding 1798 [incl. M. eurystomus Swainson 1833, Centronotus eurystomus]RD75
`--M. fulvescens (Sowerby 1834)RC73
Muricidae incertae sedis:
Ocinebrellus Jousseaume 1880AV03, RD75 [=Ternaria Coen 1943RD75]
|--*O. eurypteron (Reeve 1845) [=Murex eurypteron, *Ternaria eurypteron]RD75
|--O. aduncusAV03
`--O. nagaokaiAV03
Uttleya Marwick 1934P66
|--*U. arcana Marwick 1934RD75
|--U. ahiparana (Powell 1927)P61
|--U. marwicki Powell 1952P61
`--U. williamsi Powell 1952P66
Ceratostoma Hermannsen 1846 non Fr. 1818 (ICBN)AV03, RD75 (see below for synonymy)
|--*C. nuttallii (Conrad 1837) [=Murex (*Cerastoma) nuttallii, *Spinostoma nuttalli]RD75
|--C. burnettiAV03 [=Cerastoma foliatum var. burnettiiC64]
|--C. foliatum (Martyn 1784)AV03, O27 (see below for synonymy)
|--C. inornatumCH97
`--C. monocerosC64
Xymene Iredale 1915P61 [=Kalydon Hutton 1884 (nom. rej.)RD75]
|--*X. plebeia (Hutton 1873)RD75 [=Fusus plebeiusRD75, Trophon (*Kalydon) plebejusF26, RD75]
|--‘Ricinula’ adelaidensis [=Kalydon adelaidensis]H08
|--X. ambiguusHS01
|--X. drewi (Hutton 1883) [=Cominella drewi]F26
|--X. expansa (Hutton 1883) [=Trophon expansus]F26
|--X. inferus (Hutton 1873)P61 [=Fusus inferusF26, Trophon (Kalydon) inferusF26]
|--‘Purpura’ neglecta [=Kalydon neglecta]H08
|--X. pulcherrima (Suter 1917) [=Trophon (Xanthochorus) pulcherrimus]F26
`--X. traversiHS01
ZeatrophonP61
Xymenella Finlay 1927P61
|--*X. pusilla (Suter 1907)RD75 [=Trophon (Trophonopsis) pusillusF26]
|--X. asperula Powell & Bartrum 1929PB29
|--X. gouldi (Cossmann 1903) [=Trophon gouldi, Xymene oliveri Marwick 1924]F27
|--X. inambitiosa Marwick 1929PB29
|--X. lepida (Suter 1917) [=Trophon (Kalydon) lepidus]F26
|--X. minutissima (Suter 1917) [=Trophon (Kalydon) minutissimus]F26
`--X. suteri (Marshall & Murdoch 1921) [=Cymatium suteri]F26
Paratrophon Finlay 1927P61
|--*P. cheesemani (Hutton 1882)RD75 [=Polytropa cheesemaniRD75, Trophon (Xanthochorus) cheesemaniF26]
|--P. exsculptus Powell 1933P61
|--P. patens (Hombron & Jacquinot 1854)P61
|--P. quoyiHS01
`--P. stangeri (Gray 1843)P61 (see below for synonymy)
AxymeneP61
Comptella Finlay 1927P61
|--*C. curta (Murdoch 1905)RD75 [=Trophon (Kalydon) curtaD56, T. (K.) curtusF26]
|--C. coronata Dell 1956P61
`--C. devia (Suter 1908)P61 [=Mangilia deviaF26]
TerefundusP61
Vesanula Finlay 1926F26
|--*V. chaskanon (Finlay 1926) [=Trophon chaskanon]RD75
|--V. maorum (M. & M. 1923) [=Columbarium maorum]F26
|--V. tegens (Hutton 1877)F26
`--V. vegrandis (M. & M. 1923) [=Pagodula vegrandis]F26
Hadriania Bucquoy & Dautzenberg 1882F27, RD75
|--H. craticuloides Vokes 1964 (see below for synonymy)RD75
`--H. basedowi Cossmann 1903 [=Murex irregularis Tate 1888 non Bellardi 1873]F27
Urosalpinx Stimpson 1865O27
|--*U. cinereus (Say 1822) [=Fusus cinereus]RD75
| |--U. c. cinereusD88
| `--U. c. follyensis Baker 1951D88
|--U. macra Verrill 1884R72
|--U. perrugata (Conrad 1846)RC73
`--U. tampaensis (Conrad 1846)RC73
Eupleura Adams & Adams 1853O27
|--*E. caudata (Say 1822) [=Ranella caudata]RD75
| |--E. c. caudataD88
| `--E. c. etterae Baker 1951D88
|--E. grippi Dall 1911O27
|--E. muriciformis (Broderip 1833)KSD71
`--E. sulcidentataRC73
Forreria Jousseaume 1880O27
`--*F. belcheri (Hinds 1844)RD75 [=Murex belcheriRD75, Chorus belcheriO27, M. (Hemifusus) belcheriC64]
Brocchitas Finlay 1927DK08
`--*B. exilis (Tate 1888)DK08 (see below for synonymy)
Pseudosalpinx Olsson, Harbison et al. 1953RD75
`--*P. floridana (Conrad 1837)RD75 [=Urosalpinx floridanaO27; incl. Murex ostrearum Conrad 1846RD75]
Rugotyphis Vella 1961RD75
`--*R. francescae (Finlay 1924)RD75 [=Typhis francescaeF26]
Pagodula Monterosato 1884MG-H11, RD75 [Pagodulinae]
|--*P. echinata (Kiener 1840) [=Fusus echinatus]BR17
|--P. carduelis (Watson 1886)MG-H11
`--P. vaginata (Cristofori & Jan 1832) [=Murex vaginata]RD75
Bizetiella Radwin & D’Attilio 1972BC01, RD75
|--*B. carmen (Lowe 1935) [=Tritonalia carmen]RD75
`--B. shaskyi Radwin & D’Attilio 1972BC01
Calotrophon Hertlein & Strong 1951BC01, RD75
|--‘Tritonalia’ turrita Dall 1919 [incl. *Calotrophon bristolae Hertlein & Strong 1951]RD75
|--C. andrewsi Vokes 1976BC01
`--C. ostrearum (Conrad 1846)RC73
Distichotyphis Keen & Campbell 1964BC01
`--*D. vemae Keen & Campbell 1964BC01
Haustellotyphis Jousseaume 1880BC01, RD75
|--*H. cumingi (Broderip 1833) [=Typhis cumingi]RD75
`--H. wendita Hertz 1995BC01
Ocinebrina Jousseaum 1880BC01, RD75 (see below for synonymy)
|--O. aciculata (Lamarck 1822)D88 (see below for synonymy)
`--O. adansoni Kosuge & Fernandes 1989BC01
PoirieriaBC01
Daphnellopsis Schepman 1913BK11, P66
|--*D. lamellosa Schepman 1913P66
`--D. murex Hedley 1922P66
Concepcionella Bandel & Stinnesbeck 2000BD12
`--*C. bonillana Bandel & Stinnesbeck 2000BD12
PterymarchiaBD12
|--P. martinetanaFG19
`--P. tripterus (Born 1778)BD12
Haustrum Perry 1811BR05 [=Lepsia Hutton 1884RD75; HaustrinaeBR05]
|--*H. zelandicum Perry 1811BR17
|--H. baileyana (Tenison Woods 1881) [=Neothais baileyana]W94
|--H. dentex Perry 1811C64
|--H. haustorium (Gmelin 1791)P61 (see below for synonymy)
`--H. pictum Perry 1811C64
Tripterotyphis Pilsbry & Lowe 1932 [Tripterotyphinae]BR05
|--*T. lowei (Pilsbry 1931)RD75 [=Typhis loweiRD75, Pterotyphis (Tripterotyphis) loweiPP78]
| |--T. l. loweiW94
| |--T. l. colemani (Ponder 1972)W94
| `--T. l. norfolkensis (Fleming 1962)W94
|--T. arcana (DuShane 1969) [=Pterotyphis (Tripterotyphis) arcana]BC01
|--T. cancellata (Sowerby 1841) [=Typhis (Tripterotyphis) cancellatus]W64
|--T. fayae (Keen & Campbell 1964) [=Pterotyphis (Tripterotyphis) fayae]BC01
`--T. robustus (Verco 1895)W94
TyphinaeW64
MuricopsinaeDM88
|--MuricopsisBR05
|--FavartiaDM88
|--MurexiellaBC01
|--MurexsulBC01
`--Homalocantha Mörch 1852W94
|--*H. scorpio (Linnaeus 1758)RD75 (see below for synonymy)
|--H. anatomica (Perry 1811) [incl. H. pele Pilsbry 1921, H. rota Mawe 1823]WG71
|--H. secunda (Lamarck 1822) [incl. H. lamberti Poirier 1883]W94
`--H. zamboi (Burch & Burch 1960) [=Murex (Homalocantha) anatomica zamboi]BC01
‘Nassarina’ grayiR77
Chathamidia Dell 1956D56
`--*C. expeditionis Dell 1956RD75
Shaskyus Burch & Campbell 1963V77, RD75
`--*S. festivus (Hinds 1844)RD75 [=Murex (Triremis) festivusRD75, O27]
CoralliophilinaeBR17
Aspella Mörch 1877BR05 [AspellinaeBR17]
|--*A. anceps (Lamarck 1822) [=Ranella anceps]RD75
|--A. paupercula (Adams 1850)ED65
|--A. platylaevis Radwin & D’Attilio 1976W94
|--A. ponderi Radwin & D’Attilio 1976W94
`--A. producta (Pease 1861)W94
Maxwellia Bailey 1950JB12, RD75
|--*M. gemma (Sowerby 1879)RD75 [=Murex gemmaRD75, Mu. (Triremis) gemmaO27]
`--M. angermeyerae (Emerson & D’Attilio 1965) [=Aspella (Favartia) angermeyerae]BC01
Neorapana Cooke 1918JB12, RD75
`--*N. muricata (Broderip 1832) [=Purpura muricata]RD75
Scalaspira Conrad 1862P12, RD75
`--*S. strumosa (Conrad 1862) [=Fusus strumosa]RD75
Lirosoma Conrad 1862P12
Chesatrophon Petuch 1988P12
Chesathais Petuch 1988P12
`--C. lindae Petuch 1988P12
Trisecphora patuxentia (Petuch 1988)P12
Mariasalpinx Petuch 1988P12
Patuxentrophon Petuch 1992P12
Stephanosalpinx Petuch 1988P12
`--S. candelabra Petuch 1988P12
Laevityphis Cossmann 1903P12, RD75
|--‘Typhis’ muticus Sowerby 1834 [incl. T. coronarius Deshayes 1865, *Laevityphis coronarius]RD75
`--L. acuticosta (Conrad 1830)P12
Ecphorosycon kalyx Petuch 1988P12
Phyllonotus Swainson 1833BC01, RD75
|--P. margaritensis (Abbott 1958)B59 (see below for synonymy)
|--P. bicolorC64
|--P. brassicaC64
|--P. eversoni D’Attilio, Myers & Shasky 1987BC01
|--P. nigritusC64
|--P. oculatus (Reeve 1845)V90 [=Murex oculatusW62, Chicoreus (Phyllonotus) oculatusV90]
|--P. pomum (Gmelin 1791)V90 (see below for synonymy)
`--P. princepsC64
VokesimurexPMB15
|--V. dolichourus (Ponder & Vokes 1988)WBS-S15
`--V. lindajoyceae (Petuch 1987)PMB15
Indothais lacera (Born 1778)WBS-S15
Latiaxena ficula (Reeve 1848)WBS-S15
Reishia bitubercularis (Lamarck 1822)WBS-S15
Thalessa Adams & Adams 1853WBS-S15, RD75
|--*T. hippocastanum (Gmelin 1791) [=Murex hippocastanum, Purpura hypocastaneum (l. c.)]RD75
`--T. aculeata (Deshayes 1844)WBS-S15
Timbellus de Gregorio 1885WBS-S15, RD75
|--*T. latifolius (Bellardi 1872)RD75
`--T. bednalli (Brazier 1877)WBS-S15
Leptotrophon surprisensisFG19
Bedevina Habe 1946RC73, RD75
`--*B. birileffi (Lischke 1871) [=Trophon birileffi]RD75
*Boreotrophon clathrata (Linné 1767)RD75 [=Murex clathrataRD75, Tritonium (Trophon) clathratumC64, Trophon clathratusN79; incl. Tro. costatumJ64b, Tritonium gunneri Lovén 1846N79, Fusus gunneriC64, Trophon clathratus var. gunneriN79, F. peruvianusJ64b, Murex rumphiusJ64b, F. scalariformisJ64b, Trophon scalariformisJ64a]
Brocchitas exilis (Tate 1888)DK08 [=Fasciolaria exilisF27, Fusus exilis nec Menke 1843 nec Philippi 1843 (not preoc. if in dif. gen.)F27, Latirofusus exilisF27, Streptochetus exilisF27, S. adelomorphus Cossmann 1901F27]
Ceratostoma Hermannsen 1846 non Fr. 1818 (ICBN)AV03, RD75 [=Cerastoma Conrad 1837 non Latreille 1802RD75, Spinostoma Coen 1943RD75]
Ceratostoma foliatum (Martyn 1784)AV03, O27 [=Purpura foliataO27, Murex foliatusO27; incl. Cerostoma monodonC64, M. phyllopterusC64, M. tripterusC64]
Drupella ochrostoma (Blainville 1832)RA75 [=Drupa (Drupella) ochrostomaRA75, Purpura ochrostomaH09, Sistrum ochrostomaC64]
Hadriania craticuloides Vokes 1964 [incl. Murex craticulatus Brocchi 1814 non Linnaeus 1758, *Hadriania craticulata]RD75
Haustrum haustorium (Gmelin 1791)P61 [=Buccinum haustoriumRD75; incl. B. haustrum Martyn 1788 (nom. inv.)RD75, Haustrum haustrumRD75, *Lepsia haustrumRD75, Thais haustrumF26]
Hexaplex Swainson 1833W94, BC01 [incl. Bassia Jousseaume 1880 nec All. 1766 (ICBN) nec Quoy & Gaimard 1830RA75, Bassiella Wenz 1941W94]
Hexaplex stainforthi (Reeve 1843)W94 [=Murex stainforthiRA75, *Bassia stainforthiRA75, *Bassiella stainforthiRA75]
*Homalocantha scorpio (Linnaeus 1758)RD75 [=Murex scorpioRD75; incl. H. digitatus Sowerby 1841W94, H. fauroti Jousseaume 1888W94, H. varicosus Sowerby 1834W94]
*Naquetia triqueter (Born 1778)W94 [=Murex triqueter RA75, Pterynotus (*Naquetia) triqueterWG71; incl. N. cancellata Röding 1798W94, N. flexuosa Perry 1811W94, N. roseotinctus Sowerby 1860W94, N. variegata Röding 1798W94]
Naquetia cumingii (Adams 1853) [incl. N. triqueter var. amanuensis Couturier 1907, N. trigonulus Lamarck 1822 non Lamarck 1816]W94
Ocinebrina Jousseaum 1880BC01, RD75 [incl. Corallinia Buzquoy & Dautzenberg in Bucquoy, Dautzenberg & Dollfus 1882RD75]
Ocinebrina aciculata (Lamarck 1822)D88 [=Murex aciculatusRD75, *Corallinia aciculataRD75; incl. Fusus corallinus Scacchi 1836RD75, *Ocinebrina corallinaRD75]
Paratrophon stangeri (Gray 1843)P61 [=Fusus stangeriF26, Trophon stangeriF26, T. rugosus (Quoy & Gaimard 1833) (preoc.)F26; incl. Polytropa retiaria Hutton 1878F26]
Phyllonotus margaritensis (Abbott 1958)B59 [=Murex (Phyllonotus) margaritensisB59, M. imperialis Swainson 1831 non Fischer de Waldheim 1807RA75, *Phyllonotus imperialisRA75]
Phyllonotus pomum (Gmelin 1791)V90 [=Murex pomumW62, Chicoreus (Phyllonotus) pomumV90; incl. M. asperrimus Lamarck 1822W62, M. globosa Emmons 1852W62, M. mexicanus Petit de la Saussaye 1852W62, M. pomiformis Mörch 1852W62]
*Type species of generic name indicated
References
[AV03] Amano, K., & G. J. Vermeij. 2003. Evolutionary adaptation and geographic spread of the Cenozoic buccinid genus Lirabuccinum in the North Pacific. Journal of Paleontology 77 (5): 863–872.
[B59] Baker, B. B. 1959. Publications received: The marine mollusks of Grand Cayman Island, British West Indies. By R. Tucker Abbott. Nautilus 72 (4): 147–148.
[BD12] Bandel, K., & D. T. Dockery III. 2012. Protoconch characters of Late Cretaceous Latrogastropoda (Neogastropoda and Neomesogastropoda) as an aid in the reconstruction of the phylogeny of the Neogastropoda. Freiberger Forschungshefte C542 (psf 20): 93–128.
Barco, A., M. Claremont, D. G. Reid, R. Houart, P. Bouchet, S. T. Williams, C. Cruaud, A. Couloux & M. Oliverio. 2010. A molecular phylogenetic framework for the Muricidae, a diverse family of carnivorous gastropods. Molecular Phylogenetics and Evolution 56: 1025–1039.
[B56] Berry, S. S. 1956. Mollusca dredged by the Orca off the Santa Barbara Islands, California, in 1951. Journal of the Washington Academy of Sciences 46 (5): 150–157.
[BK11] Bouchet, P., Y. I. Kantor, A. Sysoev & N. Puillandre. 2011. A new operational classification of the Conoidea (Gastropoda). Journal of Molluscan Studies 77: 273–308.
[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.
[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.
[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.
[BW09] Bryce, C., & C. Whisson. 2009. The macromolluscs of Mermaid (Rowley Shoals), Scott and Seringapatam Reefs, Western Australia. Records of the Western Australian Museum Supplement 77: 177–208.
[C64] Carpenter, P. P. 1864. Supplementary report on the present state of our knowledge with regard to the Mollusca of the west coast of North America. Report of the British Association for the Advancement of Science 33: 517–686.
[CH97] Castro, P., & M. E. Huber. 1997. Marine Biology 2nd ed. WCB McGraw-Hill: Boston.
[DK08] Darragh, T. A., & G. W. Kendrick. 2008. Silicified Eocene molluscs from the Lower Murchison district, Southern Carnarvon Basin, Western Australia. Records of the Western Australian Museum 24 (3): 217–246.
[D88] D’Asaro, C. N. 1988. Micromorphology of neogastropod egg capsules. Nautilus 102 (4): 134–148.
[DM84] D’Attilio, A., & B. W. Myers. 1984. Descriptions of five new muricacean gastropods and comments on two additional species, in the families Muricidae and Coralliophilidae: (Mollusca). Transactions of the San Diego Society of Natural History 20 (5): 81–94.
[DM88] D’Attilio, A., & B. W. Myers. 1988. A new species of Favartia from the eastern Pacific (Gastropoda: Muricidae). Nautilus 102 (3): 106–109.
[D56] Dell, R. K. 1956. The archibenthal Mollusca of New Zealand. Dominion Museum Bulletin 18: 1–235.
[ED65] Emerson, W. K., & A. D’Attilio. 1965. Aspella (Favartia) angermeyerae, n. sp. Nautilus 79 (1): 1–4.
[FG19] Fedosov, A. E., M. C. Gutierrez, B. Buge, P. V. Sorokin, N. Puillandre & P. Bouchet. 2019. Mapping the missing branch on the neogastropod tree of life: molecular phylogeny of marginelliform gastropods. Journal of Molluscan Studies 85: 439–451.
[F26] Finlay, H. J. 1926. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.
[F27] Finlay, H. J. 1927. New specific names for austral Mollusca. Transactions and Proceedings of the New Zealand Institute 57: 488–533.
[GAS03] Gulbin, V. V., I. S. Arzamastsev & V. M. Shulkin. 2003. Ecological monitoring of the water area of Port Vostochnyi (Wrangel Bay) in the Sea of Japan (1995–2002). Russian Journal of Marine Biology 29 (5): 284–295.
[HS01] Hayward, B. W., A. B. Stephenson, M. S. Morley, W. M. Blom, H. R. Grenfell, F. J. Brook, J. L. Riley, F. Thompson & J. J. Hayward. 2001. Marine biota of Parengarenga Harbour, Northland, New Zealand. Records of the Auckland Museum 37: 45–80.
[H08] Hedley, C. 1908. Studies on Australian Mollusca. Part X. Proceedings of the Linnean Society of New South Wales 33: 456–489, pls 7–10.
[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.
[J64a] Jeffreys, J. G. 1864a. Report of the committee appointed for exploring the coasts of Shetland by means of the dredge. Report of the British Association for the Advancement of Science 33: 70–81.
[J64b] Jeffreys, J. G. 1864b. The Upper Tertiary fossils at Uddevalla, in Sweden. Report of the British Association for the Advancement of Science 33 (Notices and Abstracts): 73–79.
[JB12] Johnson, M. E., & B. G. Baarli. 2012. Development of intertidal biotas through Phanerozoic time. In: Talent, J. A. (ed.) Earth and Life: Global biodiversity, extinction intervals and biogeographic perturbations through time pp. 63–128. Springer.
[KSD71] Kern, J. P., T. E. Stump & R. J. Dowlen. 1971. An Upper Pleistocene marine fauna from Mission Bay, San Diego, California. Transactions of the San Diego Society of Natural History 16 (15): 329–338.
[LB12] Luque, Á. A., A. Barrajón, J. M. Remón, D. Moreno & L. Moro. 2012. Marginella glabella (Mollusca: Gastropoda: Marginellidae): a new alien species from tropical West Africa established in southern Mediterranean Spain through a new introduction pathway. Marine Biodiversity Records 5: e17.
[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O’Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.
[N79] Norman, A. M. 1879. The Mollusca of the fiords near Bergen, Norway. Journal of Conchology 2: 8–77.
[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. 2 pt 2. Stanford University Press: Stanford University (California).
[P12] Petuch, E. J. 2012. Miocene asteroid impacts: proposed effects on the biogeography and extinction patterns of eastern North American gastropods. In: Talent, J. A. (ed.) Earth and Life: Global biodiversity, extinction intervals and biogeographic perturbations through time pp. 967–981. Springer.
[PMB15] Petuch, E. J., R. F. Myers & D. P. Berschauer. 2015. The Living and Fossil Busycon Whelks: Iconic Mollusks of Eastern North America. San Diego Shell Club, Inc.
[PP78] Poorman, F. L., & L. H. Poorman. 1978. Additional molluscan records from Bahía de Los Angeles, Baja California Norte. Veliger 20 (4): 369–374.
[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4th ed. Whitcombe and Tombs Limited: Christchurch.
[P66] Powell, A. W. B. 1966. The molluscan families Speightiidae and Turridae: an evaluation of the valid taxa, both Recent and fossil, with lists of characteristic species. Bulletin of the Auckland Institute and Museum 5: 1–184, pls 1–23.
[PB29] Powell, A. W. B., & J. A. Bartrum. 1929. The Tertiary (Waitematan) molluscan fauna of Oneroa, Waiheke Island. Transactions and Proceedings of the New Zealand Institute 60: 395–447.
[R72] Radwin, G. E. 1972. The systematic position of Urosalpinx carolinensis Verrill, 1884 with comments on the genus Mohnia Friele, 1878. Transactions of the San Diego Society of Natural History 16 (16): 339–342.
[R77] Radwin, G. E. 1977. The family Columbellidae in the western Atlantic. Part IIa.—The Pyreninae. Veliger 20 (2): 119–133.
[RC73] Radwin, G. E., & J. L. Chamberlin. 1973. Patterns of larval development in stenoglossan gastropods. Transactions of the San Diego Society of Natural History 17 (9): 107–118.
[RD75] Radwin, G. E., & A. D’Attilio. 1975. A catalogue of muricacean generic taxa. Transactions of the San Diego Society of Natural History 17 (20): 279–292.
[S11] Simone, L. R. L. 2011. Phylogeny of the Caenogastropoda (Mollusca), based on comparative morphology. Arquivos de Zoologia 42 (4): 161–323.
[V77] Vermeij, G. J. 1977. Patterns in crab claw size: the geography of crushing. Systematic Zoology 26 (2): 138–151.
[V90] Vokes, E. H. 1990. Two new species of Chicoreus subgenus Siratus (Gastropoda: Muricidae) from northeastern Brazil. Nautilus 103 (4): 124–130.
[V95] Vokes, E. H. 1995. Two additional species of Dermomurex (Gastropoda: Muricidae) from southern Australia. Molluscan Research 16: 51–58.
[W64] Warmke, G. L. 1964. A new Caribbean muricid mollusk, Typhis puertoricensis. Nautilus 78 (1): 1–3.
[W62] Weisbord, N. E. 1962. Late Cenozoic gastropods from northern Venezuela. Bulletins of American Paleontology 62: 7–672.
[WBS-S15] Willan, R. C., C. Bryce & S. M. Slack-Smith. 2015. Kimberley marine biota. Historical data: molluscs. Records of the Western Australian Museum Supplement 84: 287–343.
[W94] Wilson, B. 1994. Australian Marine Shells vol. 2. Prosobranch Gastropods. Part Two (Neogastropods). Odyssey Publishing: Kallaroo (Western Australia).
[WG71] Wilson, B. R., & K. Gillett. 1971. Australian Shells: illustrating and describing 600 species of marine gastropods found in Australian waters. A. H. & A. W. Reed: Sydney.
[ZLK11] Zou, S., Q. Li & L. Kong. 2011. Additional gene data and increased sampling give new insights into the phylogenetic relationships of Neogastropoda, within the caenogastropod phylogenetic framework. Molecular Phylogenetics and Evolution 61: 425–435.