Acanthodrilidae

Belongs within: Opisthopora.
Contains: Rhododrilus, Acanthodrilus, Eodrilus, Microscolex, Diplocardia, Megascolecidae.

The Acanthodrilidae are an assemblage of earthworms found in Australasia, southern Africa and the Americas, most of which have two pairs of prostate pores on the segments anterior and posterior to the male pore segment.

Acanthodrilidae [Acanthodrilacea, Acanthodrilinae, Dichogastrini, Eodrilacea, Maheinacea, Octochaetidae]JD12
    |--BenhamiinaeJD12
    |    |  i. s.: NeogasterJD12
    |    |         OmodeonaJD12
    |    |         PickfordiaJD12
    |    |         Wegeneriella divergensJD12, R84b
    |    |           |--W. d. divergensR84b
    |    |           `--W. d. itapecu Righi et al. 1978R84b
    |    |         WegenerionaJD12
    |    |--BenhamionaJD12
    |    `--+--MillsoniaJD12
    |       `--DichogasterJD12
    |            |--D. affinis (Michaelsen 1890) [=Benhamia affinis]R84c
    |            |--D. annae (Horst 1883) [=Benhamia annae; incl. D. silvestris cacaois Righi 1968]R84a
    |            |--D. bolaui (Michaelsen 1891) [=Benhamia bolaui]R84c
    |            |--D. damonisM13
    |            |--D. frickeiM13
    |            |--D. gracilis (Michaelsen 1892) (see below for synonymy)R84c
    |            |--D. modiglianii (Rosa 1896) [=Benhamia modiglianii]R96
    |            |--D. nyassanaM13
    |            |--D. oraedividitis (Cognetti 1904)J88
    |            |--D. reinekeiM13
    |            |--D. saliens (Beddard 1892) [=Microdrilus saliens]R96b
    |            |--D. servi Righi & Ayres 1975R84b
    |            |--D. taboranaM13
    |            `--D. utipanaM13
    `--+--+--Parachilota Pickford 1937JD12, J12
       |  `--DiplotremaJD12
       |       |--*D. fragilis Spencer 1900M13, J88 [=Plutellus (Diplotrema) fragilisM13]
       |       |--D. acropetraB07
       |       |--D. culminis (Michaelsen 1913) [=Plutellus (Diplotrema) culminis]M13
       |       |--D. glandiferaB07
       |       |--D. ignambii (Michaelsen 1913) [=Plutellus (Diplotrema) ignambii]M13
       |       |--D. modestum (Michaelsen 1913) [=Plutellus (Diplotrema) modestus]M13
       |       |--D. rouxi (Michaelsen 1913) [=Plutellus (Diplotrema) rouxi]M13
       |       `--D. sarasini (Michaelsen 1913) [=Plutellus (Diplotrema) sarasini]M13
       `--+--Diplocardiinae [Trigastrinae]JD12
          |    |--Neotrigaster rufa (Gates 1962)JD12
          |    |--DiplocardiaJD12
          |    `--Trigaster lankesteriJD12, M13
          `--MegascolecidaeJD12

Acanthodrilidae incertae sedis:
  RhododrilusM07
  AcanthodrilusM13
  EodrilusJ31
  MicroscolexJ88
  Plagiochaeta lateralisM07
  Dinodriloides Benham 1904M07
    |--*D. beddardi Benham 1904M07
    `--D. annectensM07
  Mandane Kinberg 1866R89
    |--M. bovei Rosa 1889R89
    |--‘Acanthodrilus’ georgianusR89
    `--M. litoralis Kinberg 1866 [incl. M. patagonica Kinberg 1866]R89
  DinodrilusM07
  Octochaetus Beddard 1892JD12
    |--O. altanmouiB07
    `--O. multiporus (Beddard 1885)J88
  ZapatadrilusJD12
  ZapoteciaJD12
  ProtozapoteciaJD12
  LarsonidrilusJD12
  BalanteodrilusJD12
  RamiellonaJD12