
Belongs within: Microhylidae.
Contains: Xenorhina.
The Asterophryninae are an Indo-Australian group of frogs exhibiting direct development (Frost et al. 2006).
<==Asterophryninae [Asterophrydidae, Genyophrynidae, Genyophryninae]PW11
| i. s.: Pherohapsis Zweifel 1972FG06
|--Gastrophrynoides Moble 1926FB17, FG06
| `--G. immaculatusFB17
`--+--+--+--+--‘Liophryne’ schlaginhaufeniFB17
| | | `--Genyophryne Boulenger 1890PW11, FG06
| | | `--G. thomsoniPW11
| | `--Copiula Mehely 1901PW11, DE08
| | |--C. majorPW11
| | `--+--+--C. obstiPW11
| | | `--C. pipiensPW11
| | `--Austrochaperina Fry 1912PW11, FG06
| | |--A. adelphe Zweifel 1985C18
| | |--A. derongoPW11
| | |--A. fryi Zweifel 1962C18
| | |--A. gracilipes (Fry 1912)C18
| | |--A. pluvialis Zweifel 1965C18
| | `--A. robusta (Fry 1912)C18
| `--+--+--+--Liophryne Boulenger 1897PW11, FG06
| | | | `--L. rhododactylaPW11
| | | `--Oxydactyla Kampen 1913PW11, FG06
| | | `--O. crassaPW11
| | `--+--‘Xenobatrachus’ obesus Zweifel 1960FG06 [=Xenorhina obesaPW11]
| | |--‘Liophryne’ dentataPW11
| | `--+--‘Hylophorbus’ rufescensPW11
| | `--Sphenophryne Peters & Doria 1878PW11, FG06 [Sphenophryninae]
| | |--S. cornutaPW11
| | `--S. gracilipesAR99
| `--+--+--‘Callulops’ robustusPW11
| | `--Hylophorbus Macleay 1878PW11, FG06
| | |--H. nigrinusPW11
| | `--+--H. wondiwoiPW11
| | `--+--H. picoidesPW11
| | `--H. tetraphonusPW11
| `--+--‘Barygenys’ flavigularisPW11
| `--+--XenorhinaPW11
| `--+--Mantophryne Boulenger 1897FB17, FG06
| | `--M. lateralisFB17
| `--Callulops Boulenger 1888PW11, FG06 [Callulopini]
| | i. s.: C. wilhelmanusFB17
| |--+--C. eurydactylusPW11
| | `--C. pulliferPW11
| `--+--C. slateriPW11
| `--Asterophrys Tschudi 1838PW11, FG06
| `--A. turpicolaPW11
`--+--+--+--‘Cophixalus’ sphagnicolaPW11
| | `--Barygenys Parker 1936PW11, FG06 [Barygenini]
| | `--B. exsulPW11
| `--+--Choerophryne Van Kampen 1914PW11, DE08
| | `--C. rostelliferPW11
| `--Albericus Burton & Zweifel 1995PW11, FG06
| `--A. lauriniPW11
`--+--Aphantophryne Fry 1917PW11, FG06
| `--A. pansaPW11
`--+--Oreophryne Boettger 1895PW11, FG06
| |--+--O. clamataPW11
| | `--O. wairaPW11
| `--+--+--+--O. atrigularisPW11
| | | `--O. wapogaPW11
| | `--+--O. brachypusPW11
| | `--O. unicolorPW11
| `--+--O. sibilansPW11
| `--+--‘Cophixalus’ balbusPW11
| `--+--O. asplenicolaPW11
| `--O. pseudasplenicolaPW11
`--Cophixalus Boettger 1892FB17, FG06
|--C. aenigma Hoskin 2004C18
|--C. australis Hoskin 2012C18
|--C. bombiens Zweifel 1985C18
|--C. cheesmanaeFB17
|--C. concinnus Tyler 1979C18
|--C. crepitans Zweifel 1985C18
|--C. exiguus Zweifel & Parker 1969C18
|--C. hinchinbrookensis Hoskin 2012C18
|--C. hosmeri Zweifel 1985C18
|--C. humicolaPW11
|--C. infacetus Zweifel 1985C18
|--C. kulakula Hoskin & Aland 2011C18
|--C. mcdonaldi Zweifel 1985C18
|--C. monticola Richards, Dennis et al. 1994C18
|--C. neglectus Zweifel 1962C18
|--C. ornatus (Fry 1912)C18
|--C. pakayakulangun Hoskin & Aland 2011C18
|--C. peninsularis Zweifel 198C18
|--C. petrophilus Hoskin 2013C18
|--C. saxatilis Zweifel & Parker 1977C18
|--C. tridactylusPW11
`--C. zweifeli Davies & McDonald 1998C18
*Type species of generic name indicated
References
[AR99] Alford, R. A., & S. J. Richards. 1999. Global amphibian declines: a problem in applied ecology. Annual Review of Ecology and Systematics 30: 133–165.
[C18] Cogger, H. G. 2018. Reptiles and Amphibians of Australia updated 7th ed. CSIRO Publishing: Collingwood.
[DE08] Doughty, P., & D. Edwards. 2008. A new species of Arenophryne (Anura: Myobatrachidae) from the central coast of Western Australia. Records of the Western Australian Museum 24 (2): 121–131.
[FB17] Feng, Y.-J., D. C. Blackburn, D. Liang, D. M. Hillis, D. B. Wake, D. C. Cannatella & P. Zhang. 2017. Phylogenomics reveals rapid, simultaneous diversification of three major clades of Gondwanan frogs at the Cretaceous–Paleogene boundary. Proceedings of the National Academy of Sciences of the USA 114 (29): E5864–E5870.
[FG06] Frost, D. R., T. Grant, J. Faivovich, R. H. Bain, A. Haas, C. F. B. Haddad, R. O. de Sá, A. Channing, M. Wilkinson, S. C. Donnellan, C. J. Raxworthy, J. A. Campbell, B. L. Blotto, P. Moler, R. C. Drewes, R. A. Nussbaum, J. D. Lynch, D. M. Green & W. C. Wheeler. 2006. The amphibian tree of life. Bulletin of the American Museum of Natural History 297: 1–370.
[PW11] Pyron, R. A., & J. J. Wiens. 2011. A large-scale phylogeny of Amphibia including over 2800 species, and a revised classification of extant frogs, salamanders, and caecilians. Molecular Phylogenetics and Evolution 61: 543–583.