Belongs within: Plagiorchiida.
Contains: Bucephalinae.
The Bucephalata are a clade of flukes that exhibit a sporocyst stage in bivalve molluscs (Olson et al. 2003).
<==Bucephalata
|--Bucephalidae [Bucephaloidea, Gasterostomata, Gasterostomidae]OC03
| |--BucephalinaeD56
| `--ProsorhynchinaeD56
| |--Niedhartia Nagaty 1937 [incl. Pseudoprosorhynchus Yamaguti 1938]D56
| | |--N. ghardagae Nagaty 1937D56
| | `--N. niedharti Nagaty 1937D56
| |--Alcicornis MacCallum 1917D56
| | |--A. baylisi Nagaty 1937D56
| | `--A. carangis MacCallum 1917D56
| `--Prosorhynchus Odhner 1905 (see below for synonymy)D56
| |--*P. crucibulum (Rudolphi 1819) (see below for synonymy)D56
| |--P. aculeatus Odhner 1905 [=Skrjabiniella aculeatus]D56
| |--P. arabiana Srivastava 1937D56
| |--P. caudovatus Manter 1940D56
| |--P. facilis (Ozaki 1924) [=Gotonius facilis; incl. P. apertus McFarlane 1936]D56
| |--P. freitasi Nagaty 1937D56
| |--P. gonoderus Manter 1940D56
| |--P. manteri Srivastava 1937D56
| |--P. ozakii Manter 1934D56
| |--P. pacificus Manter 1940D56
| |--P. platycephali (Yamaguti 1934) [=Gotonius platycephali]D56
| |--P. rotundus Manter 1940D56
| |--P. squamatus Odhner 1905 (see below for synonymy)D56
| |--P. uniporus Ozaki 1924 [=Skrjabiniella uniporus]D56
| `--P. vaneyi Shen 1930 [=Dollfusina vaneyi, Dollfustrema vaneyi]D56
`--GymnophalloideaOC03
| i. s.: Gymnophallus Odhner 1905 [Gymnophallinae]D56
| |--G. affinis Jameson & Nicoll 1913D56
| |--G. bursicola Odhner 1900D56
| |--G. choledochus Odhner 1900D56
| |--G. diapsilis Nicoll 1907D56
| |--G. deliciosus (Olsson 1893) [=Distoma deliciosum]D56
| |--G. macroporus Jameson & Nicoll 1913D56
| |--G. micropharyngeus (Lühe 1898)D56
| |--G. oidemiae Jameson & Nicoll 1913D56
| |--G. ovoplenus Jameson & Nicoll 1913D56
| |--G. somateriae (Levinsen 1881) (see below for synonymy)O05
| `--‘Cercaria’ strigata Lebour 1908D56
|--Prosogonarium [Tandanicolidae]OC03
| `--P. angelaeOC03
`--Fellodistomidae [Fellodistomatinae, Steringophoridae]OC03
| i. s.: Steringotrema Odhner 1911D56
| |--S. cluthense (Nicoll 1909) [=Steringophorus cluthensis, Leioderma cluthense]D56
| |--S. divergens (Rudolphi 1809) [=Distoma divergens; incl. Fasciola blennii Müller 1776]D56
| `--S. pagelli (Beneden 1870) (see below for synonymy)D56
| Rhodotrema Odhner 1911D56
| |--R. ovacutum (Lebour 1908) [=Steringophorus ovacutus]D56
| |--R. problematicum Issaitschikow 1928D56
| |--R. quinquelobata Layman 1930D56
| `--R. skrjabini Issaitschikow 1928D56
| Ovotrema Pigulewsky 1938D56
| `--*O. ponticaD56
| Choanomyzus tasmaniae Manter & Crowcroft 1950BC02
|--Steringophorus Odhner 1905OC03, D56 [=Leioderma Stafford 1904 nec Suhm 1873 nec Nyl. 1888 (ICBN)D56]
| |--*S. furciger (Olsson 1868)O05, D56 [=Distoma furcigerumD56, Distomum furcigerumD56, *Leioderma furcigerumO05]
| `--S. margolisiOC03
`--+--+--Olssonium turneriOC03
| `--Fellodistomum Stafford 1904OC03, D56
| |--*F. fellis (Olsson 1868)O05, D56 (see below for synonymy)
| |--F. furcigerumC02
| `--F. melanostigmumK-MC02
`--HaplocladinaeD56
|--Ancyclocoelium Nicoll 1912D56
| `--A. typicum Nicoll 1912D56
|--Tergestia Stossich 1899 [incl. Theledera Linton 1910]D56
| |--T. acanthocephala (Stossich 1887)D56
| |--T. clonacantha Manter 1963MK00
| `--T. laticollis (Rudolphi 1819) (see below for synonymy)D56
|--Proctoeces Odhner 1911OC03, D56
| |--P. maculatus (Looss 1901) [=Distomum maculatum; incl. P. erythraeus Odhner 1911]D56
| `--P. magnorus Manter 1940D56
`--Haplocladus Odhner 1911D56
|--H. filiformis (Rudolphi 1819)D56
|--H. minor Odhner 1911D56
`--H. typicus Odhner 1911D56
*Fellodistomum fellis (Olsson 1868)O05, D56 [=Distoma fellisD56; incl. F. agnotum Nicoll 1909D56, Adolescaria ophiurae Tauson 1917BK77]
Gymnophallus somateriae (Levinsen 1881) [=Distomum somateriae, Lecithodendrium somateriae, Levcithodendrium (l. c.) somateriae]O05
Prosorhynchus Odhner 1905 [incl. Dollfusina Eckmann 1932 (preoc.), Dollfustrema Eckmann 1934, Gotonius Ozaki 1924, Mordvilkovia Pigulewsky 1931, Skrjabiniella Issaitschikow 1928]D56
*Prosorhynchus crucibulum (Rudolphi 1819) [=Monostomum crucibulum, Gasterostomum crucibulum; incl. G. armatum Molin 1859, Prosorhynchus costai Travassos, Artigas & Pereira 1928, Mordvilkovia elongata Pigulewsky 1931, P. scalpellus McFarlane 1936]D56
Prosorhynchus squamatus Odhner 1905 [incl. Bucephalus crux Levinsen 1881, P. grandis Lebour 1908, P. triglae Nicoll 1914]D56
Steringotrema pagelli (Beneden 1870) [=Distoma pagelli; incl. Distomum actaeonis Pagenstecher 1863, Distomum tergestinum Stossich 1889]D56
Tergestia laticollis (Rudolphi 1819) [=Distoma laticolle; incl. Cercaria dichotoma Muller in La Valette 1855]D56
*Type species of generic name indicated
References
[BK77] Barel, C. D. N., & P. G. N. Kramers. 1977. A survey of the echinoderm associates of the north-east Atlantic area. Zoologische Verhandelingen 156: 1–159.
[BC02] Bray, R. A., & T. H. Cribb. 2002. Observations on the phylogeny of Opisthadena Linton, 1910 and related genera (Hemiuridae: Opisthadeninae) from Australian and French Polynesian waters. Folia Parasitologica 49: 279–290.
[C02] Collette, B. B. 2002. Wrymouths. Family Cryptacanthodidae. In: Collette, B. B., & G. Klein-MacPhee (eds) Bigelow and Schroeder’s Fishes of the Gulf of Maine 3rd ed. pp. 483–485. Smithsonian Institute Press: Washington.
[D56] Dawes, B. 1956. The Trematoda with special reference to British and other European forms 2nd ed. University Press: Cambridge.
[K-MC02] Klein-MacPhee, G., & B. B. Collette. 2002. Eelpouts. Family Zoarcidae. In: Collette, B. B., & G. Klein-MacPhee (eds) Bigelow and Schroeder’s Fishes of the Gulf of Maine 3rd ed. pp. 466–474. Smithsonian Institute Press: Washington.
[MK00] Machida, M., & T. Kuramochi. 2000. Digenean trematodes from halfbeaks and needlefishes of Japan and adjacent waters. Bulletin of the Natural Science Museum Series A (Zoology) 26: 203–218.
[O05] Odhner, T. 1905. Die Trematoden des arktischen Gebietes. In: Römer, F., & F. Schaudinn (eds) Fauna Arctica. Eine Zusammenstellun der arktischen Tierformen, mit besonder Berücksichtigung des Spitzbergen-Gebietes auf Grund der Ergebnisse der Deutschen Expedition in das Nördliche Eismeer im Jahre 1898 vol. 4 pp. 289–372. Gustav Fischer: Jena.
[OC03] Olson, P. D., T. H. Cribb, V. V. Tkach, R. A. Bray & D. T. J. Littlewood. 2003. Phylogeny and classification of the Digenea (Platyhelminthes: Trematoda). International Journal for Parasitology 33: 733–755.