Belongs within: Caryophyllales.
Contains: Anacampseros section Avonia, Anacampseros section Telephiastrum, Didiereoideae, Portulaca, Claytonia, Calandrinia, Calyptridium, Lewisia, Cactoideae, Opuntioideae.
The Cactineae are a clade of flowering plants strongly supported by molecular analyses. Among its members are the cacti (Cactaceae), the purslanes (Portulacaceae) and the Madagascar-endemic Didereaceae. Many members of the clade are succulent to some degree. Members of the mostly Neotropical Cactaceae are spiny and most except the basal genus Pereskia lack permanent leaves.
Other members of the clade include the Basellaceae, a tropical and subtropical group of perennial herbs, many of them climbers growing from tuberous rhizomes. The Hectorellaceae are dense cushion-forming perennial herbs found in southern New Zealand and Kerguelen. Anacampseros is an African genus of succulents with alternate, usually closely placed leaves. Montia is a genus of more or less fleshy herbs bearing capsular fruits with three valves whose margins roll inwards to forcibly expel seeds (Hickman 1993).
<==Cactineae [Portulaceae]
| i. s.: Talinaria palmeriGP09
|--BasellaceaeAW03
| |--Basella rubraPT01
| |--Anredera cordifolia [incl. Boussingaultia gracilis var. pseudobaselloides]H93
| |--Ullucus tuberosusH93, S00
| `--Boussingaultia gracilisM68
|--Hectorellaceae [Pycnophylleae]T00
| |--LyalliaT00
| |--PycnophyllumA61
| `--Hectorella Hook. f. 1864A61
| `--H. caespitosa Hook. f. 1864 [incl. H. elongata Buchan. 1884]A61
|--AnacampserosCV06 [AnacampserotaceaeAPG16]
| | i. s.: A. polyphylla [=Rulingia polyphylla, Talinum polyphyllum]J54
| |--A. sect. AvoniaJ54
| |--A. (sect. Rosulatae) dielsianaJ54
| |--A. sect. TelephiastrumJ54
| `--A. (sect. Tuberosae) australianaH90
|--DidiereaceaeAW03
| |--+--Calyptrotheca Gilg 1897 [Calyptrothecoideae]AW03
| | `--DidiereoideaeAW03
| `--Portulacarioideae [Portulacarieae]AW03
| |--Portulacaria Jacquin 1787AW03
| | |--P. afraO98
| | `--P. armianaGP09
| `--Ceraria Pearson & Stephens 1912AW03
| |--C. longipedunculataCV06
| |--C. namaquensis [=Portulaca namaquensis]J54
| `--C. schaeferiJ54
|--Montia Linnaeus 1753A61 [MontiaceaeAPG16]
| |--M. chamissoiH93 [=Claytonia chamissoiM68]
| |--M. dichotomaH93
| |--M. diffusaH93
| |--M. fontana Linnaeus 1753 [incl. M. lamprosperma Cham. 1831]G95
| | |--M. f. ssp. fontanaH90
| | |--M. f. ssp. amporitana [incl. M. fontana ssp. intermedia]H90
| | `--M. f. ssp. chondrospermaH90
| |--M. funstoniiM68
| |--M. halliiM68
| |--M. howelliiH93
| |--M. linearisH93
| |--M. minor [incl. M. verna]M68
| `--M. parvifoliaH93 [=Claytonia parvifoliaM68]
|--PortulacaceaeAW03
| |--Neopaxia australasicaME70 (see below for synonymy)
| |--TalinellaG04
| |--GrahamiaG04
| |--TalinopsisG04
| |--PortulacaG95
| |--ClaytoniaA61
| |--CalandriniaC06
| |--CalyptridiumH93
| |--LewisiaH93
| |--Rumicastrum chamaecladum (Diels) Ulbr. 1934 [=Atriplex chamaeclada Diels 1919]W84
| `--Talinum [Talinaceae]G95
| |--T. paniculatum (Jacq.) Gaertn. 1791 (see below for synonymy)G95
| `--T. triangulare (Jacq.) Willdenow in Linnaeus 1799 (see below for synonymy)G95
`--CactaceaeAW03
| i. s.: MammillariaH93
| |--M. dioica [incl. M. dioica var. incerta]H93
| |--M. leucocentraPT01
| |--M. milleriH93
| `--M. tetrancistraH93
| Epiphyllum oxypetalumP88
| PilosocereusH11
| |--P. arrabidaeH11
| |--P. gounelleiBV09
| |--P. sartorianusI92
| |--P. tillianusK06
| `--P. tuberculatusBV09
| Stenocereus griseusK06
| Subpilocereus repandusK06
| Schlumbergera × buckleyiMH98
| Lophocereus schottiGE05
| EchinopsisH06
| |--E. multiplexT84
| `--E. spachianaH06
| EchinocactusH93
| |--E. polycephalusH93
| `--E. wislizenniLW70
| Carnegiea gigantea [=Cereus giganteus]B38
| Bergerocactus emoryi [=Cereus emoryi]H93
| EchinocereusH93
| |--E. brandegeeiB71
| |--E. engelmannii [incl. E. munzii]H93
| `--E. triglochidiatus [incl. E. mojavensis]H93
| Escobaria vivipara [=Coryphantha vivipara, Mammillaria vivipara]H93
| |--E. v. var. viviparaH93
| |--E. v. var. alversonii [=Coryphantha vivipara var. alversonii, Mammillaria vivipara var. alversonii]H93
| |--E. v. var. deserti [=Coryphantha deserti, Mammillaria deserti]H93
| `--E. v. var. rosea [=Coryphantha vivipara var. rosea]H93
| FerocactusH93
| |--F. cylindraceus [=Echinocactus cylindraceus]H93
| | |--F. c. var. cylindraceusH93
| | `--F. c. var. leconteiH93
| |--F. johnstonianusM69
| |--F. viridescensH93
| `--F. wislizeniiSBM11
| SclerocactusH93
| |--S. johnsonii [=Echinocactus johnsonii, Neolloydia johnsonii]H93
| `--S. polyancistrus [=Echinocactus polyancistrus]H93
| Phyllocactus akkermanniS00
| Eopuntia douglassii Chaney 1944CBH93
| Hariotia Adans. 1763KC01
| Cleistocactus burmanniiT84
|--Maihuenia [Maihuenioideae]GP09
| |--M. patagonicaGP09
| `--M. poeppigiiGP09
`--+--CactoideaeG04
`--+--OpuntioideaeG04
`--Pereskia Miller 1754GP09, T84 [incl. RhodocactusG04; Pereskieae, Pereskioideae]
|--*P. aculeata Miller 1754T84
|--P. diaz-roemerianaG04
|--P. grandifoliaG04
|--P. humboldtiiG04
`--P. weberianaG04
Neopaxia australasicaME70 [=Claytonia australasica Hook. f. in Hook. 1840H90, A61, Montia australasica (Hook. f.) Pax & Hoffman 1934A61, Paxia australasicaH90; incl. C. australasica var. biflora Buchanan 1871A61, C. calycina Colenso 1896A61, Montia calycina (Colenso) Pax & Hoffman 1934A61, C. australasica var. racemosa Buchanan 1871A61, C. australasica var. sessiliflora Simpson 1952A61]
Talinum paniculatum (Jacq.) Gaertn. 1791 [=Portulaca paniculatum Jacq. 1760; incl. T. patens (Linnaeus) Willdenow 1799]G95
Talinum triangulare (Jacq.) Willdenow in Linnaeus 1799 [=Portulaca triangulare Jacq. 1760; incl. P. racemosa Linnaeus 1771]G95
*Type species of generic name indicated
References
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