Belongs within: Cottiodei.
Contains: Cottinae, Psychrolutidae.
The Cottoidea, sculpins, are marine and freshwater fishes with large spiny heads, wide gill openings, broad mouths, slender bodies, large fan-like pectoral fins, small caudal fins, and pelvic fins reduced to three long rays (Collette & Klein-MacPhee 2002). They include the Agonidae, poachers, which are cold-water marine fishes with the body covered with bony plates.
Cottoidea [Icelidae]
|--+--CottidaeB-RW17
| | |--Leptocottus armatusB-RW17
| | `--CottinaeND13
| `--PsychrolutidaeB-RW17
`--+--ScorpaenichthysND13 [ScorpaenichthyidaeB-RW17]
| `--S. marmoratus Ayres 1854ND13, CWH77
`--Agonidae [Hemitripteridae]B-RW17
| i. s.: Leptagonus decagonus Bloch & Schneider 1801K-M02c
| Agonopsis chiloensis (Jenyns 1840)LD09
| Agonus cataphractusLM13
| PodothecusP93
| |--P. costulatus Stinton 1978P93
| `--P. sachiUT08
| Hemitripterus [Hemitripteridae]K-M02b
| `--H. americanus (Gmelin 1789)K-M02b
|--HemilepidotusB-RW17
| |--H. hemilepidotusN85
| |--H. jordaniB-RW17
| `--H. zapusB-RW17
`--+--Hypsagonus quadricornisB-RW17
`--+--SarritorB-RW17
| |--S. frenatusB-RW17
| `--S. leptorhynchusB-RW17
`--+--AspidophoroidesB-RW17
| |--A. monopterygius (Bloch 1786)K-M02c
| `--A. olrikiD56
`--+--Stellerina xyosternaB-RW17
`--BathyagonusB-RW17
|--B. alascanusB-RW17
`--+--B. pentacanthusB-RW17
`--Xeneretmus latifrons (Gilbert 1890)B-RW17, LD09
Cottoidea incertae sedis:
IcelusK-M02a
Taurulus bubalis Euphrasen 1786LD09
Trachydermus [incl. Centridermichthys]JR10
`--T. fasciatus Heckel 1840 [=Trachidermus fasciatus; incl. Trachydermus ansatus]JR10
Eocottus veronensis (Volta 1796)P93 [incl. Gobius macrurus Agassiz 1833–1844B80]
AntipodocottusN85
|--A. elegans Fricke & Brunken 1984N85
|--A. galatheaeN85
`--A. megalops DeWitt 1969N85
StlengisN85
|--*S. osensisN85
|--S. distoechusN85
|--S. mesembrinus Fricke & Brunken 1983N85
`--S. misakiaN85
Atopocottus Bolin 1936N85
`--A. tribranchiusN85
Ascelichthys rhodorusN85
ClinocottusN85
|--C. acuticepsN85
|--C. analisB96
`--C. globicepsB96
Jordania [Jordaniidae]N85
`--J. zonopeN85
Oligocottus maculosusN85
Alcichthys alcicornisN85
Elaphichthys elongatusN85
Phasmatocottus ctenopterygiusN85
PseudoblenniusN85
*Type species of generic name indicated
References
[B-RW17] Betancur-R, R., E. O. Wiley, G. Arratia, A. Acero, N. Bailly, M. Miya, G. Lecointre & G. Ortí. 2017. Phylogenetic classification of bony fishes. BMC Evolutionary Biology 17: 162.
[B80] Blot, J. 1980. La faune ichthyologique des gisements du Monte Bolca (Province de Vérone, Italie). Catalogue systématique présentant l’état actuel des recherches concernant cette faune. Bulletin du Museum National d’Histoire Naturelle C 2: 339–396.
[B96] Bond, C. E. 1996. Biology of Fishes 2nd ed. Saunders College Publishing: Fort Worth.
[CWH77] Cooper, J., M. Wieland & A. Hines. 1977. Subtidal abalone populations in an area inhabited by sea otters. Veliger 20 (2): 163–167.
[D56] Dawes, B. 1956. The Trematoda with special reference to British and other European forms. University Press: Cambridge.
[JR10] Jordan, D. S., & R. E. Richardson. 1910. Check-list of the species of fishes known from the Philippine archipelago. Bureau of Printing: Manila.
[K-M02a] Klein-MacPhee, G. 2002a. Sculpins. Family Cottidae. In: Collette, B. B., & G. Klein-MacPhee (eds) Bigelow and Schroeder’s Fishes of the Gulf of Maine 3rd ed. pp. 346–357. Smithsonian Institute Press: Washington.
[K-M02b] Klein-MacPhee, G. 2002b. Sea ravens. Family Hemitripteridae. In: Collette, B. B., & G. Klein-MacPhee (eds) Bigelow and Schroeder’s Fishes of the Gulf of Maine 3rd ed. pp. 358–361. Smithsonian Institute Press: Washington.
[K-M02c] Klein-MacPhee, G. 2002c. Alligatorfishes. Family Agonidae. In: Collette, B. B., & G. Klein-MacPhee (eds) Bigelow and Schroeder’s Fishes of the Gulf of Maine 3rd ed. pp. 361–362. Smithsonian Institute Press: Washington.
[LM13] Lautredou, A.-C., H. Motomura, C. Gallut, C. Ozouf-Costaz, C. Cruaud, G. Lecointre & A. Dettai. 2013. New nuclear markers and exploration of the relationships among Serraniformes (Acanthomorpha, Teleostei): the importance of working at multiple scales. Molecular Phylogenetics and Evolution 67: 140–155.
[LD09] Li, B., A. Dettaï, C. Cruaud, A. Couloux, M. Desoutter-Meniger & G. Lecointre. 2009. RNF213, a new nuclear marker for acanthomorph phylogeny. Molecular Phylogenetics and Evolution 50: 345–363.
[ND13] Near, T. J., A. Dornburg, R. I. Eytan, B. P. Keck, W. L. Smith, K. L. Kuhn, J. A. Moore, S. A. Price, F. T. Burbrink, M. Friedman & P. C. Wainwright. 2013. Phylogeny and tempo of diversification in the superradiation of spiny-rayed fishes. Proceedings of the National Academy of Sciences of the USA 110 (31): 12738–12743.
[N85] Nelson, J. S. 1985. On the relationship of the New Zealand marine fish Antipodocottus galatheae with the Japanese Stlengis misakia (Scorpaenifores: Cottidae). NZOI Records 5 (1): 1–12.
[P93] Patterson, C. 1993. Osteichthyes: Teleostei. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 621–656. Chapman & Hall: London.
[UT08] Urakawa, H., Y. Tajima, Y. Numata & S. Tsuneda. 2008. Low temperature decreases the phylogenetic diversity of ammonia-oxidizing archaea and bacteria in aquarium biofiltration systems. Applied and Environmental Microbiology 74 (3): 894–900.