Drupa

Belongs within: Rapaninae.

Drupa is an Indo-Pacific genus of whelks with multi-pointed side cusps on the radular teeth (Wilson 1994).

Characters (Wilson 1994): Shells medium-sized, thick and spinose, low-spired, ventral side flat; anterior canal short, narrow, slightly upturned, posterior canal represented by sulcus across ventral side of outer lip below most posterior spine; aperture heavily, elaborately denticulate on both sides, labial teeth typically grouped but sometimes evenly spaced apart. Central teeth of radula tricuspid, side cusps bifid to quadrifid, outer margins denticulate; laterals simple, L-shaped.

<==Drupa Röding 1798BR05 (see below for synonymy)
    |--D. (Drupa)W94
    |    |--*D. (D.) morum Röding 1798RD75 (see below for synonymy)
    |    `--D. (D.) ricina (Linnaeus 1758)W94, BD12 (see below for synonymy)
    |           |--D. r. ricinaBC01
    |           `--D. r. hadari Emerson & Cernohorsky 1973BC01
    `--D. (Ricinella Schumacher 1817)W94
         |--D. (R.) rubusidaeus Röding 1798W94 (see below for synonymy)
         `--D. (R.) clathrata (Lamarck 1798)W94

Drupa incertae sedis:
  ‘Sistrum’ affine Pease 1863K65
  ‘Ricinula’ arachnoideaBS29
  D. cavernosa (Reeve 1846) [=Ricinula cavernosa]H09
  D. chaidea (Duclos 1832) [=Purpura chaidea]H09
  D. concatenata (Blainville 1832) [=Purpura concatenata]H09
  D. didyma Schwengel 1943W62
  ‘Ricinula’ elegans Broderip & Sowerby 1829BS29
  D. grossularia (Röding 1798)BW09 (see below for synonymy)
  D. heptagonalis (Reeve 1846) [=Ricinula heptagonalis]H09
  ‘Sistrum’ indigoferum Melville 1901M01
  ‘Ricinula’ lugubrisG79
  D. marginalba (Blainville 1832) [=Purpura marginalba]H09
  D. muricata (Reeve 1846) [=Ricinula muricata]H09
  D. ozenneana (Crosse 1861) [=Ricinula ozenneana]H09
  D. porphyrostoma (Reeve 1846) [=Ricinula porphyrostoma]H09
  D. spinosa (Adams & Adams 1853) [=Pentadactylus spinosus]H09
  ‘Ricinula’ turbinellaG79 [incl. R. carbonariaG79, Sistrum carbonariumC64, R. ferruginosaG79]

Drupa Röding 1798^BR05 [incl. Canrena Link 1807^W94, Pentadactylus Klein in Bruguière 1792 non Schultze 1760^RD75, Ricinula Lamarck 1816^W94, Sistrum Montfort 1810^W94; Drupinae]

*Drupa (Drupa) morum Röding 1798^RD75 [incl. D. globosa Martyn in Mörch 1852^W94, *Ricinula horida Lamarck 1816^W94, *Canrena neritoidea Link 1807^W94, Drupa nodosa Linnaeus 1758^W94, D. violacea Schumacher 1817^W94]

Drupa (Drupa) ricina (Linnaeus 1758)^{W94, BD12} [=Murex ricinus^RD75, *Pentadactylus ricinus^RD75; incl. Drupa albolabris Blainville 1832^W94, *Sistrum album Montfort 1810^{W94, RD75}, D. arachnoides Lamarck 1816^WG71, D. hystrix Linnaeus 1758^W94, D. rubuscestus Dall 1915^W94, D. tribulus Röding 1798^W94]

Drupa (Ricinella) rubusidaeus Röding 1798^W94 [incl. D. fragum Röding 1798^W94, D. hippocastanum Wood 1825^W94, D. hystrix Link 1807 non Linnaeus 1758^W94, D. miticula Lamarck 1822^W94, D. nodus Gmelin 1791^W94, *Ricinella purpurata Schumacher 1817^{W94, RD75}, D. reeveana Crosse 1862^W94, D. spathulifera Blainville 1832^W94]

*Type species of generic name indicated

References

[BD12] Bandel, K., & D. T. Dockery III. 2012. Protoconch characters of Late Cretaceous Latrogastropoda (Neogastropoda and Neomesogastropoda) as an aid in the reconstruction of the phylogeny of the Neogastropoda. Freiberger Forschungshefte C542 (psf 20): 93–128.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[BS29] Broderip, W. J., & G. B. Sowerby. 1829. Observations on new or interesting Mollusca contained, for the most part, in the Museum of the Zoological Society. Zoological Journal 4: 359–379, pl. 9.

[BW09] Bryce, C., & C. Whisson. 2009. The macromolluscs of Mermaid (Rowley Shoals), Scott and Seringapatam Reefs, Western Australia. Records of the Western Australian Museum Supplement 77: 177-208.

[C64] Carpenter, P. P. 1864. Supplementary report on the present state of our knowledge with regard to the Mollusca of the west coast of North America. Report of the British Association for the Advancement of Science 33: 517–686.

[G79] Guppy, R. J. L. 1879. First sketch of a marine invertebrate fauna of the Gulf of Paria and its neighbourhood. Part I.—Mollusca. Journal of Conchology 2: 151–172.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[K65] Kay, E. A. 1965. Marine molluscs in the Cuming collection, British Museum (Natural History) described by William Harper Pease. Bulletin of the British Museum (Natural History): Zoology Supplement 1: 1–96, 14 pls.

[M01] Melville, J. C. 1901. A few further remarks upon the Erythrean molluscan fauna, with descriptions of seven species from Aden, in the collection of Commander E. R. Shopland, R.I.M. Annals and Magazine of Natural History, series 7, 7: 550–556.

[RD75] Radwin, G. E., & A. D’Attilio. 1975. A catalogue of muricacean generic taxa. Transactions of the San Diego Society of Natural History 17 (20): 279–292.

[W62] Weisbord, N. E. 1962. Late Cenozoic gastropods from northern Venezuela. Bulletins of American Paleontology 62: 7–672.

[W94] Wilson, B. 1994. Australian Marine Shells vol. 2. Prosobranch Gastropods. Part Two (Neogastropods). Odyssey Publishing: Kallaroo (Western Australia).

[WG71] Wilson, B. R., & K. Gillett. 1971. Australian Shells: illustrating and describing 600 species of marine gastropods found in Australian waters. A. H. & A. W. Reed: Sydney.