Belongs within: Globigerinana.
The Heterohelicidae are a group of planktonic foraminiferans known from the Late Cretaceous to the Oligocene. The test is planispiral or biserial in early stages and may develope chamber proliferation or serial reduction in later stages (Loeblich & Tappan 1964).
Characters (Loeblich & Tappan 1964, as Heterohelicinae): Early stage planispiral or biserial, lateral may show chamber proliferation or serial reduction; aperture simple and interiomarginal, or terminal in uniserial forms.
Heterohelicidae
|--Gublerina Kikoïne 1948LT64 [incl. Sigalia Reiss 1957LT64; GublerininaeM13]
| |--G. ornatissima (Cushman & Church 1929) [=Ventilabrella ornatissima; incl. *G. cuvillieri Kikoïne 1948]LT64
| `--G. deflaensis (Sigal 1952) [=Guembelina (Ventilabrella) deflaensis, *Sigalia deflaensis]LT64
|--Pseudoguembelina Brönnimann & Brown 1953LT64 [PseudogumbelininaeM13]
| `--*P. excolata (Cushman 1926) [=Guembelina excolata]LT64
`--Heterohelicinae [Guembelininae]M13
|--Racemiguembelina Montanaro Gallitelli 1957LT64
| `--*R. fructicosa (Egger 1899) [=Guembelina fructicosa]LT64
|--Bifarina Parker & Jones 1872 [incl. Rectoguembelina Cushman 1932, Tubitextularia Šulc 1929]LT64
| |--*B. saxipara (Ehrenberg 1854) [=Dimorphina saxipara]LT64
| |--B. bohemica [=Pseudotextularia bohemica, *Tubitextularia bohemica]LT64
| |--B. fimbriataC40
| |--B. nodosaria White 1929 [incl. *Rectoguembelina cretacea Cushman 1932]LT64
| `--‘Rectoguembelina’ texanaC40
|--Heterohelix Ehrenberg 1843 [=Spiroplecta Ehrenberg 1844; incl. Guembelina Egger 1899 non Kuntze 1895]LT64
| |--*H. americana (Ehrenberg 1844) [=*Spiroplecta americana; incl. H. navarroensis]LT64
| |--H. cenomanicaBL79
| |--H. complanataM08b
| |--H. dentata Stenestad 1968FM03
| |--H. globulosa (Ehrenberg 1840) [=Textularia globulosa, *Guembelina globulosa]LT64
| |--H. moremaniI02
| |--H. pulchraM08b
| |--‘Guembelina’ spiniferaC40
| `--H. striataM08b [=Guembelina striataC40]
|--Planoglobulina Cushman 1927 [incl. Ventilabrella Cushman 1928]LT64
| |--*P. acervulinoides (Egger 1899) [=Guembelina acervulinoides]LT64
| |--‘Ventilabrella’ carseyiC40
| `--‘*Ventilabrella’ eggeri Cushman 1928LT64
`--Pseudotextularia Rzehak 1891 (see below for synonymy)LT64
|--*P. elegans (Rzehak 1891) [=Cuneolina elegans]LT64
|--P. fruticosaC40
|--‘Guembelina’ plummerae Loetterle 1937 [=*Bronnimannella plummerae]LT64
`--P. varians Rzehak 1886C40
Pseudotextularia Rzehak 1891 [incl. Bronnibrownia Montanaro Gallitelli 1955 (n. n.), Bronnimannella Montanaro Gallitelli 1956]LT64
*Type species of generic name indicated
References
[BL79] Basov, V. A., B. G. Lopatin, I. S. Gramberg, A. I. Danjushevskaya, V. Ya. Kaban’kov, V. M. Lazurkin & D. K. Patrunov. 1979. Lower Cretaceous lithostratigraphy near Galicia Bank. Initial Reports of the Deep Sea Drilling Project 47: 683–717.
[C40] Cushman, J. A. 1940. Foraminifera: Their classification and economic use 3rd ed. Harvard University Press: Cambridge (Massachusetts).
[FM03] Friedrich, O., & K. J. S. Meier. 2003. Stable isotope indication for the cyst formation depth of Campanian/Maastrichtian calcareous dinoflagellates. Micropaleontology 49 (4): 375–380.
[I02] Ibrahim, M. I. A. 2002. Late Albian–Middle Cenomanian palynofacies and palynostratigraphy, Abu Gharadig-5 well, Western Desert, Egypt. Cretaceous Research 23: 775–788.
[LT64] Loeblich, A. R., Jr & H. Tappan. 1964. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 2. The Geological Society of America and The University of Kansas Press.
[M08] McMillan, I. K. 2008. Reappraisal of foraminiferal assemblages of the Santonian–Campanian Mzamba Formation type section, and their correlation with the stratigraphic succession of the KwaZulu Basin. African Natural History 4: 25–34.
[M13] Mikhalevich, V. I. 2013. New insight into the systematics and evolution of the Foraminifera. Micropaleontology 59 (6): 493–527.