Hyaenidae

Striped hyaena Hyaena hyaena, copyright Balaji Venkatesh Sivaramakrishnan.

Belongs within: Aeluroidea.

The Hyaenidae, hyaenas, are cursorial carnivores known from Africa and Eurasia, with fossil representatives in North America. Hyaenas have relatively short bodies with conspicuously down-sloping backs, and are conspicuously marked with stripes or spots.

<==Hyaenidae
    |--CrocutaFS15
    |    |--C. brevirostrisWC56
    |    |--C. crocutaRB84
    |    |    |--C. c. crocutaRB84
    |    |    `--C. c. spelaeaRB84
    |    |--C. eximiaK54
    |    |--C. honanensisDW04
    |    `--C. variabilisK54
    `--+--Proteles cristatus (Sparrman 1783)FS15, V10
       `--HyaenaFS15
            |--H. abroniaS78
            |--H. bellaxS78
            |--H. brunneaS78
            |--H. crocutaG41
            |--H. hyaena (Linnaeus 1758)DC02
            |    |--H. h. hyaenaUSDI77
            |    `--H. h. barbaraUSDI77
            |--H. makapani Toerien 1952WC56
            |--H. priscusME05
            |--H. spelaeaF82
            |--H. striataK54
            |--H. ultimaJ63
            `--H. vulgarisG41

Hyaenidae incertae sedis:
  Parahyaena brunneaYC04
  Hyaenictis preforfexS78
  Leecyaena forfex [=Hyaenictis forfex]S78
  ProgenettaS78
  Miohyaena certaK54
  ChasmaporthetesYC04
    |--C. ossifragusV91
    `--C. progressusDW04
  EuryboasS78
    |--E. namaquensis [=Hyaena namaquensis, Lycyaena namaquensis]S78
    `--E. silberbergi [=Lycyaena silberbergi]S78
  PachycrocutaME05
    |--P. brevirrostrisWT08
    |--P. licentiDW04
    |--P. robustaJ63
    `--P. sinensisDW04
  Percrocuta Schlosser 1903S78, D07
    |--P. algeriensis [=Hyaena algeriensis]S78
    |--P. australis Hendey 1974S78
    |--P. giganteaD07
    |--P. hebiensisD07
    |--P. miocenicaD07
    |--P. tobieni Crusafont & Aguirre 1971S78
    `--P. tungurensisD07
  Ictitherium Wagner 1848 (see below for synonymy)K54
    |  i. s.: I. arkesilaiD07
    |         I. ebuWT08
    |         I. ibericumD07
    |         I. intuberculatumD07
    |         I. kurteniD07
    |         I. orbignyiK54
    |         I. pannonicumD07
    |--+--I. hipparionum [=Hyaena hipparionum]K54
    |  `--I. wongiiK54 [=Hyaenictitherium wongiiDW04]
    `--+--I. robustum (Nordmann ex Gervais 1858) (see below for synonymy)K54
       |    |--I. r. robustumK54
       |    `--I. r. tauricumK54
       `--+--I. gaudryiK54
          `--+--I. hyaenoidesK54 [=Hyaenictitherium hyaenoidesDW04]
             |--I. indicumK54
             `--I. sivalenseK54
  Herpestides de Beaumont 1967SM93
  ProtictitheriumWT08
  TungurictisWT08
  PlioviverropsWT08
  Adcrocuta variabilisWT08, DW04
Inorganic: Proteles cristatus minilorientalus Okamura 1987O87

Ictitherium Wagner 1848 [incl. Galeotherium Wagner 1840 non Jäger 1839, Hyaenictitherium Kretzoi 1937, Lepthyaena Lydekker 1884, Palhyaena Gervais 1859, Sinictitherium Kretzoi 1937, Thalassictis Nordmann ex Gervais 1850]K54

Ictitherium robustum (Nordmann ex Gervais 1858) [=Thalassictis robusta; incl. T. gracilis Hensel 1862, Ictitherium viverrinum Roth & Wagner 1854]K54

*Type species of generic name indicated

References

[DC02] De, J. K., & R. Chakraborty. 2002. Identification of dorsal guard hairs of striped hyena Hyaena hyaena (Linnaeus, 1758) Hyaenidae: Carnivora: Mammalia. Journal of the Bombay Natural History Society 99 (3): 502–506.

[DW04] Deng T., Wang X., Ni X. & Liu L. 2004. Sequence of the Cenozoic mammalian faunas of the Linxia Basin in Gansu, China. Acta Geologica Sinica (English Edition) 78 (1): 8–14.

[D07] Dixon, D. 2007. The Complete Illustrated Encyclopedia of Dinosaurs & Prehistoric Creatures. Hermes House: London.

[FS15] Faurby, S., & J.-C. Svenning. 2015. A species-level phylogeny of all extant and late Quaternary extinct mammals using a novel heuristic-hierarchical Bayesian approach. Molecular Phylogenetics and Evolution 84: 14–26.

[F82] Finch, M. E. 1982. The discovery and interpretation of Thylacoleo carnifex (Thylacoleonidae, Marsupialia). In: Archer, M. (ed.) Carnivorous Marsupials vol. 2 pp. 537–551. Royal Zoological Society of New South Wales.

[G41] Gulliver, G. 1841. Observations on the blood-corpuscles of the order Ferae. Proceedings of the Zoological Society of London 9: 42–44.

[J63] Jánossy, D. 1963. Letztinterglaziale Vertebraten-Fauna aus der Kálmán Lambrecht-Höhle (Bükk-Gebirge, nordost-Ungarn) I. Acta Zoologica Academiae Scientiarum Hungaricae 9 (3–4): 293–331.

[K54] Kurtén, B. 1954. The type collection of Ictitherium robustum (Gervais, ex Nordmann) and the radiation of the ictitheres. Acta Zoologica Fennica 86: 1–26.

[ME05] Moullé, P.-E., A. Echassoux, F. Lacombat, E. Desclaux & S. Bailon. 2005. L’environnement animal des premiers habitants de l’Europe méditerranéenne: les grands mammifères contemporains de l’homme du Vallonnet, données taxonomiques et biostratigraphiques pour la deuxième moitie du Pléistocène inférieur. BAR International Series 1364: 105–113.

[O87] Okamura, C. 1987. New facts: Homo and all Vertebrata were born simultaneously in the former Paleozoic in Japan. Original Report of the Okamura Fossil Laboratory 15: 347–573.

[RB84] Richardson, P. K. R., & S. K. Bearder. 1984. The hyaena family. In: Macdonald, D. (ed.) All the World’s Animals: Carnivores pp. 146–151. Torstar Books Inc.: New York.

[S78] Savage, R. J. G. 1978. Carnivora. In: Maglio, V. J., & H. B. S. Cooke (eds) Evolution of African Mammals pp. 249–267. Harvard University Press: Cambridge (Massachusetts).

[SM93] Stucky, R. K., & M. C. McKenna. 1993. Mammalia. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 739–771. Chapman & Hall: London.

[USDI77] United States Department of the Interior. 1977. Endangered and threatened wildlife and plants—republication of list of species. Federal Register 42: 36420–36431.

[V91] Valkenburgh, B. van. 1991. Iterative evolution of hypercarnivory in canids (Mammalia: Carnivora): evolutionary interactions among sympatric predators. Paleobiology 17 (4): 340–362.

[V10] Vršanský, P. 2010. Cockroach as the earliest eusocial animal. Acta Geologica Sinica (English Edition) 84 (4): 793–808.

[WT08] Wang, X., & R. H. Tedford. 2008. Dogs: Their fossil relatives and evolutionary history. Columbia University Press: New York.

[WC56] Wells, L. H., & H. B. S. Cooke. 1956. Fossil Bovidae from the Limeworks Quarry, Makapansgat, Potgietersrus. Palaeontologia Africana 4: 1–55.

[YC04] Yamaguchi, N., A. Cooper, L. Werdelin & D. W. Macdonald. 2004. Evolution of the mane and group-living in the lion (Panthera leo): a review. Journal of Zoology 263: 329–342.

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