
Belongs within: Aeluroidea.
The Hyaenidae, hyaenas, are cursorial carnivores known from Africa and Eurasia, with fossil representatives in North America. Hyaenas have relatively short bodies with conspicuously down-sloping backs, and are conspicuously marked with stripes or spots.
<==Hyaenidae |--CrocutaFS15 | |--C. brevirostrisWC56 | |--C. crocutaRB84 | | |--C. c. crocutaRB84 | | `--C. c. spelaeaRB84 | |--C. eximiaK54 | |--C. honanensisDW04 | `--C. variabilisK54 `--+--Proteles cristatus (Sparrman 1783)FS15, V10 `--HyaenaFS15 |--H. abroniaS78 |--H. bellaxS78 |--H. brunneaS78 |--H. crocutaG41 |--H. hyaena (Linnaeus 1758)DC02 | |--H. h. hyaenaUSDI77 | `--H. h. barbaraUSDI77 |--H. makapani Toerien 1952WC56 |--H. priscusME05 |--H. spelaeaF82 |--H. striataK54 |--H. ultimaJ63 `--H. vulgarisG41 Hyaenidae incertae sedis: Parahyaena brunneaYC04 Hyaenictis preforfexS78 Leecyaena forfex [=Hyaenictis forfex]S78 ProgenettaS78 Miohyaena certaK54 ChasmaporthetesYC04 |--C. ossifragusV91 `--C. progressusDW04 EuryboasS78 |--E. namaquensis [=Hyaena namaquensis, Lycyaena namaquensis]S78 `--E. silberbergi [=Lycyaena silberbergi]S78 PachycrocutaME05 |--P. brevirrostrisWT08 |--P. licentiDW04 |--P. robustaJ63 `--P. sinensisDW04 Percrocuta Schlosser 1903S78, D07 |--P. algeriensis [=Hyaena algeriensis]S78 |--P. australis Hendey 1974S78 |--P. giganteaD07 |--P. hebiensisD07 |--P. miocenicaD07 |--P. tobieni Crusafont & Aguirre 1971S78 `--P. tungurensisD07 Ictitherium Wagner 1848 (see below for synonymy)K54 | i. s.: I. arkesilaiD07 | I. ebuWT08 | I. ibericumD07 | I. intuberculatumD07 | I. kurteniD07 | I. orbignyiK54 | I. pannonicumD07 |--+--I. hipparionum [=Hyaena hipparionum]K54 | `--I. wongiiK54 [=Hyaenictitherium wongiiDW04] `--+--I. robustum (Nordmann ex Gervais 1858) (see below for synonymy)K54 | |--I. r. robustumK54 | `--I. r. tauricumK54 `--+--I. gaudryiK54 `--+--I. hyaenoidesK54 [=Hyaenictitherium hyaenoidesDW04] |--I. indicumK54 `--I. sivalenseK54 Herpestides de Beaumont 1967SM93 ProtictitheriumWT08 TungurictisWT08 PlioviverropsWT08 Adcrocuta variabilisWT08, DW04 Inorganic: Proteles cristatus minilorientalus Okamura 1987O87
Ictitherium Wagner 1848 [incl. Galeotherium Wagner 1840 non Jäger 1839, Hyaenictitherium Kretzoi 1937, Lepthyaena Lydekker 1884, Palhyaena Gervais 1859, Sinictitherium Kretzoi 1937, Thalassictis Nordmann ex Gervais 1850]K54
Ictitherium robustum (Nordmann ex Gervais 1858) [=Thalassictis robusta; incl. T. gracilis Hensel 1862, Ictitherium viverrinum Roth & Wagner 1854]K54
*Type species of generic name indicated
References
[DC02] De, J. K., & R. Chakraborty. 2002. Identification of dorsal guard hairs of striped hyena Hyaena hyaena (Linnaeus, 1758) Hyaenidae: Carnivora: Mammalia. Journal of the Bombay Natural History Society 99 (3): 502–506.
[DW04] Deng T., Wang X., Ni X. & Liu L. 2004. Sequence of the Cenozoic mammalian faunas of the Linxia Basin in Gansu, China. Acta Geologica Sinica (English Edition) 78 (1): 8–14.
[D07] Dixon, D. 2007. The Complete Illustrated Encyclopedia of Dinosaurs & Prehistoric Creatures. Hermes House: London.
[FS15] Faurby, S., & J.-C. Svenning. 2015. A species-level phylogeny of all extant and late Quaternary extinct mammals using a novel heuristic-hierarchical Bayesian approach. Molecular Phylogenetics and Evolution 84: 14–26.
[F82] Finch, M. E. 1982. The discovery and interpretation of Thylacoleo carnifex (Thylacoleonidae, Marsupialia). In: Archer, M. (ed.) Carnivorous Marsupials vol. 2 pp. 537–551. Royal Zoological Society of New South Wales.
[G41] Gulliver, G. 1841. Observations on the blood-corpuscles of the order Ferae. Proceedings of the Zoological Society of London 9: 42–44.
[J63] Jánossy, D. 1963. Letztinterglaziale Vertebraten-Fauna aus der Kálmán Lambrecht-Höhle (Bükk-Gebirge, nordost-Ungarn) I. Acta Zoologica Academiae Scientiarum Hungaricae 9 (3–4): 293–331.
[K54] Kurtén, B. 1954. The type collection of Ictitherium robustum (Gervais, ex Nordmann) and the radiation of the ictitheres. Acta Zoologica Fennica 86: 1–26.
[ME05] Moullé, P.-E., A. Echassoux, F. Lacombat, E. Desclaux & S. Bailon. 2005. L’environnement animal des premiers habitants de l’Europe méditerranéenne: les grands mammifères contemporains de l’homme du Vallonnet, données taxonomiques et biostratigraphiques pour la deuxième moitie du Pléistocène inférieur. BAR International Series 1364: 105–113.
[O87] Okamura, C. 1987. New facts: Homo and all Vertebrata were born simultaneously in the former Paleozoic in Japan. Original Report of the Okamura Fossil Laboratory 15: 347–573.
[RB84] Richardson, P. K. R., & S. K. Bearder. 1984. The hyaena family. In: Macdonald, D. (ed.) All the World’s Animals: Carnivores pp. 146–151. Torstar Books Inc.: New York.
[S78] Savage, R. J. G. 1978. Carnivora. In: Maglio, V. J., & H. B. S. Cooke (eds) Evolution of African Mammals pp. 249–267. Harvard University Press: Cambridge (Massachusetts).
[SM93] Stucky, R. K., & M. C. McKenna. 1993. Mammalia. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 739–771. Chapman & Hall: London.
[USDI77] United States Department of the Interior. 1977. Endangered and threatened wildlife and plants—republication of list of species. Federal Register 42: 36420–36431.
[V91] Valkenburgh, B. van. 1991. Iterative evolution of hypercarnivory in canids (Mammalia: Carnivora): evolutionary interactions among sympatric predators. Paleobiology 17 (4): 340–362.
[V10] Vršanský, P. 2010. Cockroach as the earliest eusocial animal. Acta Geologica Sinica (English Edition) 84 (4): 793–808.
[WT08] Wang, X., & R. H. Tedford. 2008. Dogs: Their fossil relatives and evolutionary history. Columbia University Press: New York.
[WC56] Wells, L. H., & H. B. S. Cooke. 1956. Fossil Bovidae from the Limeworks Quarry, Makapansgat, Potgietersrus. Palaeontologia Africana 4: 1–55.
[YC04] Yamaguchi, N., A. Cooper, L. Werdelin & D. W. Macdonald. 2004. Evolution of the mane and group-living in the lion (Panthera leo): a review. Journal of Zoology 263: 329–342.