Malvoideae

Belongs within: Malvaceae.
Contains: Eumalvoideae.

The Malvoideae as defined here include the traditional Malvaceae sensu stricto plus some basal taxa that were previously placed in the Bombacaceae. The clade was defined by Baum et al. (1998) as including all taxa closer to Malva sylvestris than Bombax ceiba. Possible members of this clade include the Fremontodendreae, two genera of trees and shrubs found in southern North America with a distinctive quincuncial arrangement of sepals (S. R. Hinsley). The Matisieae are a Neotropical group of trees with simple ovate or elliptical leaves, often zygomorphic flowers with a long staminal tube, and drupaceous fruits. The swizzlestick tree Quararibea turbinata produces aromatic wood that is used to flavour drinks.

Synapomorphies (from the Angiosperm Phylogeny Website): Petiole bundle annular; median calyx often abaxial, corolla basally connate to free; androecium from 5 antepetalous primordia, centrifugal, tube often with 5 apical teeth; tapetum plasmodial; pollen often spiny, 7+ porate; styles often separate, stigmas decurrent to capitate, hairy; ovules 1-many/carpel, campylotropous (rarely anatropous), outer integument 2-6 cells across, inner integument 4-8(-15) cells across, parietal tissue 2-7 cells across; embryo curved (rarely straight), cotyledons folded; n = 5-20(+).

<==Malvoideae [Metamalvaceae]
    |--FremontodendreaeBS04
    |    |--Chiranthodendron pentadactylonBS04, BAN98
    |    `--FremontodendronBS04
    |         |--F. californicumBAN98
    |         |    |--F. c. ssp. californicum (see below for synonymy)H93
    |         |    `--F. c. ssp. decumbensH93
    |         `--F. mexicanumH93
    `--+--MatisieaeBS04
       |    |--QuararibeaBS04
       |    |    |--Q. guianensisBAN98
       |    |    `--Q. turbinataSWK87
       |    `--+--Phragmotheca siderosaBS04, BAN98
       |       `--Matisia cordataBS04, BAN98
       `--+--PentaplarisBS04
          `--+--Radyera farrageiBS04, H90 [=Hibiscus farrageiH90]
             `--+--EumalvoideaeBS04
                `--+--UladendronBS04
                   `--+--CamptostemonBS04
                      |    |--C. philippinensisC81
                      |    `--C. schultzii Mast. in Hook 1872 [incl. C. aruense Becc. 1889]C81
                      `--+--Howittia trilocularisBS04, H90
                         `--Lagunaria patersoniiBS04, BAN98

Fremontodendron californicum ssp. californicum [incl. F. californicum ssp. crassifolium, F. californicum ssp. napense, F. californicum ssp. obispoense]^H93

*Type species of generic name indicated

References

[BAN98] Baum, D. A., W. S. Anderson & R. Nyffeler. 1998. A durian by any other name: taxonomy and nomenclature of the core Malvales. Harvard Papers in Botany 3 (2): 315–330.

[BS04] Baum, D. A., S. D. Smith, A. Yen, W. S. Alverson, R. Nyffeler, B. A. Whitlock & R. L. Oldham. 2004. Phylogenetic relationships of Malvatheca (Bombacoideae and Malvoideae; Malvaceae sensu lato) as inferred from plastid DNA sequences. American Journal of Botany 91: 1863–1871.

[C81] Croft, J. R. 1981. Bombacaceae. In: Henty, E. E. (ed.) Handbooks of the Flora of Papua New Guinea vol. 2 pp. 4–18. Melbourne University Press.

[H90] Harden, G. J. (ed.) 1990. Flora of New South Wales vol. 1. New South Wales University Press.

[H93] Hickman, J. C. (ed.) 1993. The Jepson Manual: Higher Plants of California. University of California Press: Berkeley (California).

[SWK87] Snyder, N. F. R., J. W. Wiley & C. B. Kepler. 1987. The Parrots of Luquillo: Natural history and conservation of the Puerto Rican parrot. Western Foundation of Vertebrate Zoology: Los Angeles.