Melitaeini

Belongs within: Nymphalinae.

The Melitaeini include the checkerspots, a primarily Nearctic group of butterflies. Checkerspots generally have wings marked with red, orange and black or brown. The genus Melitaea, fritillaries, contains about 80 species found in the Palaearctic region.

<==Melitaeini
    |--EuphydryasWWN03
    |    |--E. aniciaS72
    |    |--E. aurinia [=Melitaea aurinia]G91
    |    |--E. chalcedonaD37
    |    |--E. desfontainiiWWN03
    |    `--E. edithaS85
    |         |--E. e. edithaD37
    |         |--E. e. baroniS72
    |         `--E. e. wrightiiD37
    `--+--ChlosyneWWN03
       |    |--C. acastusS72
       |    |--C. damoetasS72
       |    |    |--C. d. damoetasS72
       |    |    `--C. d. malcolmiS72
       |    |--C. gorgoneS72
       |    |--C. hoffmanniS72
       |    |--C. ismeriaS72
       |    |--C. janais Drury 1782FS54
       |    |--C. lacinia (Geyer 1837)FS54
       |    |--C. leaniraS72
       |    |    |--C. l. leaniraS72
       |    |    `--C. l. almaS72
       |    |--C. minutaS72
       |    |--C. nycteisS72
       |    `--C. pallaS72
       `--+--MelitaeaWWN03
          |    |--M. arcesiaP01
          |    |--M. athalia [incl. M. athalia ab. navarina]P01
          |    |--M. cinxiaP01
          |    |--M. deioneOO90
          |    |--M. didymaP01 [=Argynnis (Melitaea) didymaG20; incl. M. didyma var. alpinaP01]
          |    |--M. didymoidesWWN03
          |    |--M. neumoegeniD37
          |    |--M. parthenieOO90
          |    |--M. scotosia Butler 1878I92
          |    |--M. theona (Ménétriés 1857)FS54
          |    `--M. wrightiiD37
          `--PhyciodesWWN03
               |--P. albescens Röber 1913FS54
               |--P. campestrisS72
               |--P. cocytaWWN03
               |--P. elaphiaea Hewitson 1868S55
               |--P. liriopeS55
               |    |--P. l. liriopeS55
               |    `--P. l. fragilis (Bates 1864)S55
               |--P. perillaS55
               |    |--P. p. perillaS55
               |    `--P. p. lugubris Röber 1913S55
               `--P. tharosS72

*Type species of generic name indicated

References

[D37] Dobzhansky, T. 1937. Genetics and the Origin of Species. Columbia University Press: New York.

[FS54] Franz, E., & H. Schröder. 1954. Tagfalter (Lep. Rhopalocera) aus El Salvador. Senckenbergiana Biologica 35: 75–87.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[G91] Groves, C. P. 1991. A Theory of Human and Primate Evolution revised ed. Clarendon Press: Oxford.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[OO90] Oberthür, C., & R. Oberthür. 1890. Lépidoptères dans les Pyrénées-Orientales. Annales de la Société Entomologique de France, 6^e série 9: cc–cciii.

[P01] Pável, J. 1901. Lepkék [Lepidopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 171–177. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[S55] Schröder, H. 1955. Eine Falter-Ausbeute aus dem westlichen Bolivien. (Ins. Lepid. Rhopal.) Senckenbergiana Biologica 36: 329–338.

[S72] Shields, O. 1972. Flower visitation records for butterflies (Lepidoptera). Pan-Pacific Entomologist 48 (3): 189–203.

[S85] Slatkin, M. 1985. Gene flow in natural populations. Annual Review of Ecology and Systematics 16: 393–430.

[WWN03] Wahlberg, N., E. Weingartner & S. Nylin. 2003. Towards a better understanding of the higher systematics of Nymphalidae (Lepidoptera: Papilionoidea). Molecular Phylogenetics and Evolution 28: 473–484.