Belongs within: Cimicomorpha.
Contains: Orthotylinae, Phytocoris, Phylinae, Bryocorinae, Mirinae.
The Miridae are a diverse family of true bugs including both phytophagous and carnivorous species. They can be distinguished from most other bug families by the presence of a cuneus in the forewing, together with a four-segmented labium (Carver et al. 1991). They also generally lack ocelli though these are present in the subfamily Isometopinae, sometimes treated as a distinct family.
Members of the subfamilies Cylapinae and Psallopinae possess a subapical tooth on the tarsal claws. Psallopinae have two-segmented tarsi and a single cell in the wing membrane, and lack a rounded pronotal collar. The Cylapinae have two cells in the wing membrane and usually possess three-segmented tarsi and a rounded pronotal collar, and include species believed to be mycetophagous. Members of the Mirinae, Bryocorinae, Phylinae and Orthotylinae possess pulvilli or pseudopulvilli in association with the claws. Deraeocorinae lack both subapical claw-teeth and pulvilli though the claws may possess teeth basally. The Mirinae and Deraeocorinae are united by the presence of a secondary gonopore in the male and similar genitalia in the female (Carver et al. 1991).
Characters (from Carver et al. 1991): Length 2-6 mm long; ocelli usually absent (except in Isometopinae); mesofemoral and metafemoral trichobothria present; forewing with cuneus (except in some brachypters), 1 or 2 cells in forewing membrane; abdomen without ventral trichobothria; specialised male and female genitalia.
<==Miridae [Capsidae]
|--PsallopsCGW91 [PsallopinaeW79]
`--+--PhylinaeW79
`--+--+--BryocorinaeW79
| `--MirinaeW79
`--CylapinaeW79
| i. s.: Peritropis listeri (Izzard 1936)MC13, JG19 [=Cylapofulvius listeriJG19]
|--Bakeriola [Bothriomirini]CGW91
|--Fulvius [Fulviini]CGW91
`--CylapiniCGW91
|--CarvalhomaCGW91
`--SchizopteromirisCGW91
Miridae incertae sedis:
Chaetedus reuterianusM83
HeterocordylusH01
|--H. genistaeH01
`--H. tibialisA71
Asciodema obsoletumA71
OrthotylinaeCGW91
AuricillocorisMC94
CtypomirisMC94
DioclerusMC94
MyiocapsusMC94
MertilaMC94
HarpedonaMC94
ProdromusMC94
ThaumastomirisMC94
Diomocoris russatusWFS04
Chinamiris nigrifronsWFS04
CyphodemaS54
|--C. hilareS54
|--C. instabileS54
|--C. mendosumS54
|--C. oberthuriS54
|--C. pernixS54
`--C. rubricum Seidenstücker 1954S54
DeraeocorinaeCGW91
|--Termatophyloides [Termatophylini]CGW91
|--Deraeocoris Kirschbaum 1855GM79 [incl. Estuidus Distant 1904G34a, GM79; Deraeocorini]
| |--D. brevisG01
| `--D. signatusCGW91
`--SaturniomiriniCGW91
|--Trilaccus nigroruberCGW91
`--ImogenWEE70
LygocorisYS01
MonocorisGE05
RanzoviusGE05
RagmusG34a
|--R. argenteus Girault 1934G34b
|--R. importunitasG34a
|--R. liebknechti Girault 1934G34a
|--R. pulcher Girault 1934G34a
`--R. seminigricaput Girault 1934G34b
ParacalocorisG34a
|--P. australisG34a
`--P. semiflavifemur Girault 1934G34a
Rhinomiridius bioculatus Girault 1934G34a
NesiomirisO94
Orectoderus obliquusBM76
NotostiraK08
|--N. doddiK08
|--N. elongataK09
|--N. erratica [incl. N. erratica var. ochracea]H01
`--N. pacifica Kirkaldy 1908K08
LeptopternaH01
|--L. dolabrata [incl. L. dolabrata var. aurantiaca]H01
`--L. ferrugataH01
Lopus gothicus (Linnaeus 1758)H01, L49 (see below for synonymy)
AdelphocorisH01
|--A. annulicornisH01
|--A. lineolatus [incl. A. chenopodii]H01
|--A. quadripunctatusH01
|--A. reicheliH01
|--A. seticornisH01
`--A. triannulatus [incl. A. triannulatus var. nigriceps]H01
PoeciloscytusH01
|--P. cognatusH01
|--P. unifasciatusH01
`--P. vulneratusH01
PolymerusH01
|--P. nigritaH01
`--P. pekinensis Horváth 1901H01
Camptobrochis lutescensH01
Rhopalotomus ater [incl. R. ater var. tyrannus]H01
AllodapusH01
|--A. montandoniH01
`--A. pumilus Horváth 1901H01
Systellonotus triguttatusH01
Myrmecophyes alboornatusH01
Anapus longicornisH01
Euryopocoris nitidusH01
Orthocephalus vittipennisH01
Strongylocoris leucocephalusH01
Criocoris quadrimaculatusH01
OncotylariaH01
|--OncotylusH01
|--LitoxenusH01
`--Ibiaris Horváth & Reut. in Horváth 1901H01
`--*I. discretus Horváth & Reut. in Horváth 1901H01
Isometopinae [Isometopidae]CGW91
|--Eurocrypha thanatochlamysCGW91
|--IsometopusK08
`--Letaba bedfordiRD77
Nesocrypha Kirkaldy 1908K08
`--*N. corticicola Kirkaldy 1908K08
Hyaloscytus elegantulus Reuter 1904 [incl. H. elegantulus var. filicicola Kirkaldy 1908]K08
Tichorhinus vitiensis Kirkaldy 1908K08
Nesosylphas Kirkaldy 1908K08
`--*N. pacifica Kirkaldy 1908K08
DisphinctusK08
MalacopeplusK08
CapsiniK08
|--Nesodaphne Kirkaldy 1908K08
| `--*N. knowlesi Kirkaldy 1908K08
`--CapsusE12
|--C. capillaris [=Phytocoris capillaris; incl. C. tricolor, C. vanicus]L49
|--C. minutus Lucas 1847E12
|--C. pilosusL49
`--C. rugicollis Lucas 1847E12
PhytocorisK08
TaylorilygusWEE74
|--T. aldrichi (Izzard 1936) [=Lygus aldrichi]JG19
`--T. murrayi (Izzard 1936) [=Lygus murrayi]JG19
Fushunimiris Hong 2002H02
`--*F. eocenica Hong 2002H02
‘Noliphus’ ruficollis Walker 1871D01
Cremnocephalus albolineatusR13
Blepharidopterus angulatusRD77
Plesiocoris rugicollisRD77
Sahlbergella singularisRD77
Distantiella theobromaRD77
AmbercylapusP92
ArcheofulviusP92
JordanofulviusP92
AetorhinusP92
DichrooscytusP92
HarpoceraP92
HomodemusP92
HoplomachusP92
OrthopsP92
Neurocolpus arizonaeE66
Macrolophus pygmaeusCK15
Pseudopsallus lattiniYR12
Heterotoma spissicornis [=Capsus spissicornis]L49
Lopus gothicus (Linnaeus 1758)H01, L49 [=Cimex (Seticornis) gothicusL49, L58, Capsus gothicusL49, Phytocoris gothicusL49]
*Type species of generic name indicated
References
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