Belongs within: Neritoidea.
Contains: Septaria (Navicella), Theodoxus (Theodoxus).
The Neritinidae are a family of tropical gastropods found in brackish and fresh water. Freshwater species with direct development have sometimes been distinguished as the subfamily Theodoxinae from marine forms with planktotrophic larvae (Bandel 2001).
Characters (Bandel 2001): Shell usually smooth, with very thin outer calcite layer and thick inner aragonite crossed lamellar layer composing its walls; protoconch of species with free larva convolutedly coiled and smooth or provided with fine collabral lines; inner lip of aperture forming a callus covered, flattened shelf and columellar edge commonly bearing one larger denticle on apical half and accompanying small denticles below and above, may also be smoothly straight or concave. Operculum with hinge on inner columellar side consisting of ridge that may or may not be accompanied by a peg or knob.
Neritinidae [Aspidibranchia, Neritinacea, Neritinidae, Neritininae, Neritinini, Theodoxinae, Theodoxini]BR17
|--Orthopoma Gray 1868 [Orthopomatini, Orthopomina]BR05
| `--*O. clypeolum (Récluz 1843) [=Navicella clypeolum, Septaria clypeolum]BR17
|--Stenopoma Gray 1868 [Stenopomatini, Stenopomina]BR05
| `--*S. lineata (Lamarck 1817) [=Navicella lineata]BR17
|--Fluvinerita Pilsbry 1932B01, KC60
| `--F. tenebricosa (Adams 1852)KC60 (see below for synonymy)
|--Pseudonerita Baker 1923B01
| |--*P. holosericea (Garrett 1872) [=Neritina holosericea]KC60
| `--P. obtusa (Reeve 1856)B01
|--Neritodryas Martens 1869B01
| |--*N. cornea (Linnaeus 1758) [=Nerita cornea]KC60
| |--N. chimmoi (Reeve 1856)B01
| `--N. subsulcata (Sowerby 1836)B01
|--Ninnia Brusina 1902B01 (see below for synonymy)
| |--*N. schulzei (Grimm 1877)B01 [=Neritina schultziKC60, Theodoxus (*Ninnia) schultziKC60]
| |--‘Neritaea’ colchica Andrussov 1912 [=Theodoxus (*Ninniopsis) colchicus]KC60
| |--N. geticusB01
| |--N. martensi (Brusina 1897)B01
| |--‘Neritina’ petasata Seninski 1905 [=Theodoxus (*Brusinaella) petasatus]KC60
| |--N. soceni Jekelius 1944B01
| `--‘Neritina’ unguiculatus Seninski 1905 [=Theodoxus (*Neritonyx) unguiculatus]KC60
|--Septaria Férussac 1807B01 (see below for synonymy)
| | i. s.: S. bougainvillei (Récluz 1841)B01
| | S. sanguisuga Reeve 1856B01
| | S. suffreni (Sowerby 1841)B01
| |--*S. (Septaria) borbonica (Bory de St. Vincent 1803)B01 (see below for synonymy)
| |--S. (Navicella)KC60
| |--S. (Paraseptaria Risbec 1942)KC60
| | `--S. (*P.) parva (Risbec 1942) [=*Paraseptaria parva]KC60
| `--S. (Sandalium Schumacher 1817) (see below for synonymy)KC60
| `--S. (S.) porcellana (Linnaeus 1758)KC60 (see below for synonymy)
|--Theodoxus Montfort 1810B01 (see below for synonymy)
| | i. s.: T. adelae (Brusina 1902)B01
| | T. banaticus Jekelius 1944B01
| | T. carasiensis Jekelius 1944B01
| | T. crenulatus (Klein 1853)B01
| | |--T. c. crenulatusB01
| | `--T. c. varpalotensis Bartha 1956B01
| | T. dacius Jekelius 1944B01
| | T. danasteri (Lindholm 1808)B01
| | T. dniestroviensisB01
| | T. doricus (Neumayr 1869)B01
| | T. eugenii Jekelius 1944B01
| | T. fabulus (Briart & Cornet 1887) [=Neritina fabula, T. (Vittoclithon) fabulus]B01
| | T. gizelae (Brusina 1902)B01
| | T. grateloupianus (Ferussac 1821)GK02
| | T. groyana Giroti & Esu 1974B01
| | T. intextus [=T. fluviatilis f. intexta]S63
| | T. intracarpaticus Jekelius 1944B01
| | T. liturata (Eichwald 1841) [incl. T. astrachanicus Starobogatov 1994]B01
| | T. loebersdorfensis Handmann 1887B01
| | T. maresi (Bourguignat 1864)B01
| | T. mariae (Handmann 1887)B01
| | T. matheroniB01
| | T. meridionalis (Philippi 1836)B01
| | T. micans (Fuchs 1877)B01
| | T. numidicus (Récluz 1841)B01
| | ‘Neritina’ ortoni Conrad 1871B01
| | T. perlongus Jekelius 1944B01
| | T. petralbensis Jekelius 1944B01
| | T. pitariB01
| | T. politioanei Jekelius 1944B01
| | T. politus Jekelius 1944B01
| | ‘Nerita’ pomahakaensis Finlay 1924F26 [=N. nitida (Hutton 1873) (preoc.)F27]
| | T. postcrenulatus Papp 1953B01
| | T. prozlatarici Jekelius 1944B01
| | T. sarmaticus (Lindholm 1901)B01
| | T. saulcyi (Bourguignat 1852)B01
| | T. semiplicatusB01
| | T. soceni Jekelius 1944B01
| | T. spratti (Jenkins 1864) [incl. T. abnormis (Jenkins 1864)]B01
| | T. subangularisB01
| | T. subthermalis Issel 1865B01
| | T. timisensis Jekelius 1944B01
| | T. tortuosus Jekelius 1944B01
| | T. trilophosensis Rust 1997B01
| | T. turislavicus Jekelius 1944B01
| | T. velox Anistratenko 1999B01
| | T. vetranici (Brusina 1902)B01
| | T. zlatarici (Brusina 1902)B01
| | T. zografi Brusina 1902B01
| |--T. (Theodoxus)B01
| |--T. (Calvertia Bourguignat 1880) (see below for synonymy)KC60
| | `--T. (C.) sinjanus (Brusina 1876) [=Neritina sinjana; incl. *Calvertia letournouxi Bourguignat 1880]KC60
| |--T. (Meganninia Davidachvili 1932)KC60
| | `--T. (*M.) corrugata Davidachvili 1930KC60
| `--T. (Neritaea Roth 1855)B01 [=Neritoconus Kobelt 1871KC60]
| |--T. (*N.) jordani (Sowerby 1832) (see below for synonymy)B01
| |--T. (N.) anatolicus Récluz 1841B01
| |--T. (N.) cinctilla Martens 1874B01
| |--T. (N.) euphratica Mousson 1874B01
| `--T. (N.) subterrelictus Schütt 1973BV86
`--Neritina Lamarck 1816B01 (see below for synonymy)
| i. s.: N. alata Broderip & Sowerby 1829BS29
| ‘Nerita (Clithon)’ angulosa Récluz 1842R42
| N. arctilineata Récluz in Sowerby 1849 (see below for synonymy)PB27
| N. atra Gray 1831PB27
| N. bicanaliculata (Récluz 1842) [=Nerita (Neritina) bicanaliculata]R42
| N. bicolor (Récluz 1843)B01 [=Nerita (Clithon) bicolorR42]
| ‘Clithon’ chlorostomus Broderip 1832B01
| ‘Nerita (Clithon)’ circumvoluta Récluz 1842R42
| N. consobrinaB01
| N. corona (Linnaeus 1758)B01 [=Nerita coronaB01, *Corona coronaKC60, Theodoxus (*Clithon) coronaKC60]
| N. coronata (Leach 1815) [=Clithon coronata; incl. C. longispina (Récluz 1841)]B01
| N. cuneata Gabb 1864 [=N. (Dostia) cuneata]B01
| ‘Nerita (Clithon)’ dacostae Récluz 1843R43
| ‘Clithon’ diadema (Récluz 1841)B01
| ‘Nerita (Clithon)’ donovani Récluz 1842R42
| N. dudariensis Strausz 1966B01
| N. dutempleiB01
| N. fraseri Reeve 1855PB27
| N. guayaquilensisC64
| N. intermedia [incl. N. globosa]C64
| ‘Nerita (Clithon)’ interrupta Récluz 1842R42
| N. knorri (Récluz 1841) [=Nerita knorri; incl. Neritina (Neritaea) cryptospira v. Martens 1879]PB27
| N. labiosa Sowerby 1841KC60 [=Nerita (Neritina) labiosaR42, Neritina (*Neritona) labiosaKC60]
| N. liassicaB01
| N. liturata Eichw. 1841S01
| N. lutea Zittel 1862B01
| N. melanostoma Troschel 1837 [=N. (Dostia) melanostoma]B01
| ‘Nerita’ michaudi Récluz 1841 [=N. (Clithon) michaudi]R42
| N. microstomaG79
| ‘Nerita (Clithon)’ montacuti Récluz 1842R42
| N. montensis Briant & Cornet 1887 [=Neritoplica montensis]B01
| N. natalensis Reeve 1855PB27
| N. nucleolus (Morelet 1856) [=Clithon nucleolus]B01
| ‘Nerita (Clithon)’ olivacea Récluz 1842R42 [=Clithon olivaceusB01]
| N. panayana (Récluz 1842) [=Nerita (Neritina) panayana]R42
| N. passayana Deshayes 1866B01
| N. phaseolaris White 1874W77
| N. pomahakaensis (Finlay 1924) [=Neritoplica pomahakaensis]B01
| ‘Clithon’ pritchardi (Dohrn 1861)B01
| ‘Nerita (Clithon)’ pulchella Récluz 1842R42
| N. rangiana (Recluz 1842)W93 (see below for synonymy)
| ‘Clithon’ rarispina (Mousson 1848)B01
| N. smithi (Wood 1828) [=Nerita smithi, Neritina (*Provittoida) smithi]KC60
| ‘Nerita (Clithon)’ sowerbiana Récluz 1842R42
| N. sphaerica Olsson & Harbison 1953P94
| ‘Clithon’ spinosus (Sowerby 1845)B01
| N. squamaepicta (Récluz 1842) [=Nerita (Neritina) squamaepicta]R42
| ‘Nerita (Clithon)’ squarrosa Récluz 1842R42
| N. subauriculata (Récluz 1842) [=Nerita (Neritina) subauriculata]R42
| ‘Nerita (Clithon)’ subpunctata Récluz 1843R43 [=Neritella subpunctataF66]
| N. thermophilaPB27
| N. violacea (Gmelin 1791)B01 (see below for synonymy)
| N. viridisG40
| N. zigzag (Lamarck 1822)W93 [=N. (Provittoida) zigzagW93; incl. N. coromandelianaB01]
|--+--N. cariosa (Wood 1928)B01 (see below for synonymy)
| |--N. granosa Sowerby 1825B01 [=Clypeolum granosumB01; incl. N. papillosa Jay 1836BC01]
| `--N. neglecta Pease 1860B01 [=Nerita neglectaK65, Theodoxus (*Neritoclithon) neglectusKC60]
|--+--N. communis (Quoy & Gaimard 1832) [incl. N. waigiensis]B01
| |--‘Clithon’ faba (Sowerby 1836)B01
| |--N. oualaniensis Lesson 1831B01 (see below for synonymy)
| `--N. roissyana (Wenz 1938) [=N. (*Vittina) roissyana]B01
|--+--N. gagates Lamarck 1822B01
| |--N. turrita (Gmelin 1790)B01
| |--N. turtoni (Récluz 1843)B01
| `--N. variegata (Lesson 1830)B01 [=N. (*Vittoida) variegataKC60, N. (Vittoidea) variegataB01]
|--+--N. auriculata Lamarck 1816 [=Clypeolum auriculatum, N. (Neripteron) auriculata]B01
| |--N. dilatata Broderip 1832B01
| |--N. lecontei (Récluz 1853)B01
| |--N. taitensis Lesson 1830B01 [=N. (*Neripteron) tahitiensisB01, *Neritopteron taitensisKC60]
| `--N. vespitina Sowerby 1825 [=Clypeolum vespitinum]B01
|--+--*N. pulligera (Linné 1758)B01 (see below for synonymy)
| |--N. asperulata Récluz 1855B01 [=Nerita (Neritina) asperulataR42]
| |--N. canalis Sowerby 1825B01
| |--N. macgillivrayi (Reeve 1855)B01
| |--N. petiti Récluz 1841B01 [=Nerita (Neritina) petitiR42]
| |--N. porcata Gould 1847B01
| `--N. squamipicta (Récluz 1843)B01
|--+--N. adansoniana (Récluz 1841)B01 (see below for synonymy)
| |--N. afra Sowerby 1841B01 [=Theodoxus aferPB27; incl. N. aequinoxialis Morelet 1848PB27]
| |--N. cristata Morelet 1864B01
| |--N. glabrata Sowerby 1849B01 (see below for synonymy)
| |--N. oweniana (Wood 1828)B01 (see below for synonymy)
| |--N. rubricata Morelet 1858B01 [incl. N. calabarica Mousson in v. Martens 1879PB27, N. gouldi v. Martens 1879PB27]
| `--N. tiassalensis Binder 1955B01
`--+--N. latissima Broderip 1833 [=N. (*Clypeolum) latissima, N. (Neripteron) latissima]B01
|--N. luteofasciata (Miller 1879) (see below for synonymy)B01
|--N. meleagris Lamarck 1822B01 [=Theodoxus (*Vittoclithon) meleagrisKC60]
|--N. piratica Russell 1940B01
|--N. punctulata Lamarck 1816B01 (see below for synonymy)
|--N. reclivata (Say 1822)B01
|--N. virginea (Linnaeus 1758)B01 (see below for synonymy)
`--N. zebra (Buguiere 1792)B01
Fluvinerita tenebricosa (Adams 1852)KC60 [=Neritina tenebricosaB01, Nerita tenebricosaKC60; incl. Neritina (Fluvinerita) alticola Pilsbry 1932B01, KC60, Nerita (*Fluvinerita) alticolaKC60]
Neritina Lamarck 1816B01 [=Chernites Gistel 1848KC60, Laphrostoma Rafinesque 1815KC60, LamphrostomaKC60, Lamprostoma non Swainson 1840KC60, Neritella Gray 1847BR17, Onychina Scudder 1882KC60; incl. Alina Récluz 1842 non Risso 1826KC60, Alinoclithon Baker 1923B01, Clithon Montfort 1810B01, Cliton Lesson 1830KC60, Clypeolum Récluz 1842B01, Corona Récluz 1850 non Albers 1850KC60, Dostia Gray 1847B01, Labialia Scudder 1882KC60, Mitrula Récluz 1850 nec. Pers. 1794 (ICBN) nec Fr. 1821 (ICBN) nec Gray 1821KC60, Nereina Cristofori & Jan 1832B01, Neripteron Lesson 1830B01, Neritioclithon Baker 1923B01, Neritona Martens 1869B01, Neritopteron Fischer 1885KC60, Neropterum Agassiz 1846PB27, Paranerita Bourne 1909 non Hampson 1901KC60, Pictoneritina Iredale 1936B01, Provittoida Baker 1923B01, Provittoidea Wenz 1938B01, Scapha Mörch 1852 nec Récluz 1841 nec Motschulsky 1845 nec Gray 1847KC60, Vitta Mörch 1923B01, Vittina Baker 1923B01, Vittoclithon Baker 1923B01, Vittoida Baker 1923B01, Vittoidea Wenz 1938KC60; Neritellina, Neritellinae]
Neritina adansoniana (Récluz 1841)B01 [=Nerita adansonianaPB27, Neritina (Neritaea) adansonianaPB27, Neritina (Vitta) adansonianaPB27; incl. Neritina sangara Morelet 1848PB27]
Neritina arctilineata Récluz in Sowerby 1849 [incl. N. africana Parreyss in Reeve 1856, N. dongolensis Ehrenberg in v. Martens 1865, N. nilotica Reeve 1856]PB27
Neritina cariosa (Wood 1928)B01 [=Nerita cariosusKC60, Clithon (*Alinoclithon) cariosaB01, Clypeolum cariosumB01, Theodoxus (*Alinoclithon) cariosusKC60]
Neritina glabrata Sowerby 1849B01 [=N. (Smaragdia) glabrataPB27, N.(Vitta) glabrataPB27; incl. N. glabrata var. senegalensis Duclos in Baker 1923PB27, N. webbei Récluz 1850PB27]
Neritina luteofasciata (Miller 1879) [=Theodoxus luteofaciatus, N. (Vittoclithon) luteofasciata; incl. N. picta Sowerby 1832 non Nerita picta Férussac 1825]B01
Neritina oualaniensis Lesson 1831B01 [=Clithon (*Pictoneritina) oualaniensisB01, KC60, Theodoxus (*Pictoneritina) oualaniensisW93, Vittoclithon oualaniensisW93; incl. Puperita (Heminerita) amoena (Gould 1847)B01]
Neritina oweniana (Wood 1828)B01 [=Nerita owenianaPB27, Clypeolum oweninanumB01, Neritina (Alina) owenianaPB27, Neritina (Neripteron) oweninanaB01, Neritina (Neritaea) owenianaPB27, Neritina owenii Gray 1831PB27; incl. Neritina (Neritaea) hessei Boettger 1885PB27]
*Neritina pulligera (Linné 1758)B01 [=Nerita pulligeraB01, *Chernites pulligeraKC60, Clypeolum pulligerumPB27, *Laphrostoma pulligerumKC60, *Neritella pulligeraBR17, *Onychina pulligeraKC60]
Neritina punctulata Lamarck 1816B01 [=Neritina (Nereina) punctulataB01; incl. Neritina clenchi Russell 1940B01, *Nereina lacustris Cristofori & Jan 1832KC60]
Neritina rangiana (Recluz 1842)W93 [=Nerita (Neritina) rangianaR42; incl. Nerita (Neritina) rangiana var. minor Récluz 1842R42]
Neritina violacea (Gmelin 1791)B01 [=Nerita violaceaKC60, Neritina (Dostia) violaceaB01; incl. Nerita crepidularia Lamarck 1822B01, *Mitrula crepidulariaKC60, Neritina (*Dostia) crepidulariaB01, Nerita crepidularia var. major Récluz 1842R42, Nerita mitrulaR42, Patella neritoidea Linnaeus 1758R42, L58]
Neritina virginea (Linnaeus 1758)B01 [=Nerita virgineaKC60, Neritina (*Vitta) virgineaB01, *Scapha virgineaKC60; incl. Neritina declivisG40]
Ninnia Brusina 1902B01 [incl. Brusinaela Andrussov 1912KC60, Brusinaella Andrussov 1912B01, Neritonyx Andrussov 1912B01, Ninniopsis Tomlin 1930B01]
Septaria Férussac 1807BR05 [=Cibota Adams & Adams 1858 non Herrmannsen 1852KC60, Cimber Montfort 1810KC60, Septarius Gray 1821KC60; incl. Laodia Gray 1868KC60, Paria Gray 1868 non LeConte 1858KC60; Catillina, Catillinae, Septariae, Septariidae, Septariinae, Septariini]
Septaria (Sandalium Schumacher 1817) [=Elana Gray 1867, Elara Adams & Adams 1856, Sandalinum Récluz 1841; incl. Catillus Gray 1847 non Brongniart 1822]KC60
Septaria (Sandalium) porcellana (Linnaeus 1758)KC60 [=Patella porcellanaKC60, *Catillus porcellanusBR17; incl. *Sandalium picta Schumacher 1817KC60, *Elara pictaKC60]
*Septaria (Septaria) borbonica (Bory de St. Vincent 1803)B01 [=Patella borbonicaB01, Catillus borbonicusKC60, *Cibota borbonicaKC60, *Cimber borbonicusKC60]
Theodoxus Montfort 1810B01 [=Neritoglobus Kobelt 1871KC60, TheodoraKC60, Theodorus (l. c.)KC60, Theodoxia Bourguignat 1877KC60, Theodoxis Montfort 1810KC60; incl. Elea Fitzinger 1833KC60]
Theodoxus (Calvertia Bourguignat 1880) [incl. Burgersteinia Bourguignat 1880, Petrettinia Bourguignat 1880, Saintsimonia Bourguignat 1880, Tripaloia Bourguignat 1880; incl. Neritodonta Brusina 1884]KC60
Theodoxus (*Neritaea) jordani (Sowerby 1832)B01 [=Neritina (*N.) jordaniB01, *Neritoconus jordaniKC60; incl. T. fontannesi (Neumayr 1879)B01, T. hellenicus (Bukowski 1893)B01, Neritina karasuna Mousson 1874B01, T. karasunaB01, T. macrii Récluz 1841B01, T. mesopotamica Mousson 1874B01, Neritina orontis Blanckenhorn 1897B01, T. orontisB01, T. pseudomicans (Bukowski 1893)B01]
*Type species of generic name indicated
References
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