Nuculanoidea

 Oval yoldia Yoldia myalis, photographed by L. Schroeder.

Belongs within: Pteriomorphia.
Contains: Palaeoneilo, Nuculana, Yoldia.

The Nuculanoidea are a group of infaunal bivalves whose earliest representatives are recorded from the Ordovician. Their shells are posteriorly elongate, though the degree of elongation varies from only slight to prominently so. Members of the family Nuculanidae possess a resilifer and have a partially internal ligament whereas the Malletiidae lack a resilifer and their ligament is predominantly external. The Neilonellidae resemble Malletiidae in the presence of an external ligament but have robust, ovate, concentrically ridged shells whereas Malletiidae have thin, translucent, elongate shells.

<==Nuculanoidea
    |--SiliculidaeBRW98
    |--LametilidaeBRW98
    |--MalletiidaeH00
    |    |--PalaeoneiloH00
    |    |--MalletiellaBRW98
    |    |--PseudoglomusBRW98
    |    |--NuculitesJB12
    |    |    |--N. pseudodeltoidesF71
    |    |    |--N. scissa Sherrard 1959F71
    |    |    `--N. triangula Sherrard 1959F71
    |    `--Malletia Desmoulins 1832F27a
    |         |  i. s.: M. chilensisGW02
    |         |         M. giganteaGW02
    |         |         M. inequalisGW02
    |         |         M. obtusa (Sars 1868) [=Yoldia obtusa]N79
    |         `--M. (Minormalletia Dall 1908)F27a
    |              |--M. (*M.) arciformisF27a
    |              `--M. (M.) tenera Marwick 1926F27a
    |--NeilonellidaeGW02
    |    |--Neilonella subovataGW02
    |    |--Pseudotindaria ferrari (Fleming 1950)SB93
    |    |--Protonucula verconisBRW98
    |    |--Austrotindaria Fleming 1948BRW98, P61
    |    |    |--*A. wrighti Fleming 1948P61
    |    |    |--A. benthicola Dell 1956P61
    |    |    `--A. flemingi Dell 1956P61
    |    `--Neilo Adams 1854BRW98, P61
    |         |--*N. australis (Quoy & Gaimard 1835) [=Nucula australis]P61
    |         |--N. awamoana Finlay 1926F27a
    |         |--N. cymbula (Woods 1917) [=Malletia (Neilo) cymbula]F27a
    |         |--N. ‘elongata’ (Marshall 1917) [=Malletia elongata non Etheridge 1902]F27b
    |         |--N. randsi (Etheridge 1892) [=Nuculana (Yoldia) randsi; incl. Leda elongata]F27b
    |         |--N. rugata Dell 1956P61
    |         |--N. sinangula Finlay 1926F27a
    |         |--N. sublaevis Marwick 1926F27a
    |         `--N. waitaraensis Marwick 1926F27a
    |--Nuculanidae [Yoldiidae, Yoldiinae]GW02
    |    |--SpinulaF27a
    |    |--NuculanaMG-H11
    |    |--VeteranellaBRW98
    |    |--Poroleda Hutton 1893P61
    |    |    `--*P. lanceolata (Hutton 1885) [incl. P. pertubata Iredale 1924]P61
    |    |--Zealeda Marwick 1924F27a
    |    |    |--*Z. hamata Marwick 1924F27a
    |    |    |--Z. inflatella Marwick 1931B70
    |    |    `--Z. mutabilis Marwick 1926F27a
    |    |--Thestyleda Iredale 1929P61
    |    |    |--*T. ramsayi [=Leda ramsayi]P61
    |    |    `--T. investigator Dell 1952P61
    |    |--Rollieria Cossmann 1920 (see below for synonymy)H00
    |    |    |--*R. palmae (Sowerby 1824) [=Nucula palmae]H00
    |    |    |--‘Nuculopsis’ anthraconeiloidasG31
    |    |    `--R. bronni (Andler 1858) (see below for synonymy)H00
    |    |--Ledella Verrill & Bush 1897P61
    |    |    |--*L. messanensis [=Leda messanensis]P61
    |    |    |--L. finlayi Powell 1935P61
    |    |    |--L. herdmanni Dell 1953P61
    |    |    `--L. librata Dell 1952P61
    |    `--Jupiteria Bellardi 1875P61
    |         |--*J. concava [=Nucula concava, Nuculana (*Jupiteria) concava]P61
    |         |--J. amuriensis (Woods 1917) [=Nuculana (Jupiteria) amuriensis]F27a
    |         |--J. manawatawhia (Powell 1937) [=Nuculana (Jupiteria) manawatawhia]P61
    |         |--J. onairoensis (Marwick 1926) [=Nuculana (Jupiteria) onairoensis]F27a
    |         |--J. uawaB70
    |         |--J. wolffi (Dell 1956) [=Nuculana (Jupiteria) wolffi]P61
    |         `--J. zealedaformis (Dell 1953) [=Nuculana (Jupiteria) zealedaformis]P61
    `--Sareptidae [Yoldiidae, Yoldiinae]BRW98
         |--YoldiaMG-H11
         |--MegayoldiaBRW98
         |--CnesteriumBRW98
         |--OrthoyoldiaBRW98
         |--AdrenellaBRW98
         |--MicroyoldiaBRW98
         |--YoldiellaBRW98
         |    |--Y. frigidaHG98
         |    |--Y. lenticulaHG98
         |    |--Y. lucidaGW02
         |    `--Y. nanaHG98
         |--PortlandiaBRW98
         |    |--P. arcticaHG98
         |    |--P. isonotaGW02
         |    |--P. japonica (Adams & Reeve 1850)XZ10
         |    `--P. tenuisN00
         `--Sarepta Adams 1860DK08 [incl. Ovaleda Iredale 1925BRW98, P61]
              |--S. aucklandica Marshall 1918 [=Ovoleda aucklandica]F27a
              |--‘Ovoleda’ hamiltoni Powell 1935P61
              |--S. obolella (Tate 1886) [=Leda obolella)F26
              |--S. planiuscula (Tate 1886) [=Leda planiuscula, Ovaleda planiuscula]DK08
              |--‘Ovoleda’ powelli Dell 1950P61
              `--S. tellinaeformis Hedley 1901P61, F27a [=*Ovoleda tellinaeformisF27a]

Rollieria Cossmann 1920 [incl. Isoleda Rollier 1923, Isonuculana Cox 1925, Nuculopsis Rollier 1912 non Girty 1911]H00

Rollieria bronni (Andler 1858) [=Leda bronni, Nuculana (Rollieria) bronni; incl. Tellina aequilatera Koch & Dunker 1837 (n. d.), L. aequilatera, Nucula fallax Terquem & Piette 1865, Nucula stoppani Winkler 1886]H00

*Type species of generic name indicated

References

[BRW98] Beesley, P. L., G. J. B. Ross & A. Wells (eds) 1998. Fauna of Australia vol. 5. Mollusca: The Southern Synthesis. Part A. Australian Biological Resources Study: Canberra.

[B70] Beu, A. G. 1970. Bathyal Upper Miocene Mollusca from Wairarapa district, New Zealand. Transactions of the Royal Society of New Zealand, Earth Sciences 7 (12): 209–240.

[DK08] Darragh, T. A., & G. W. Kendrick. 2008. Silicified Eocene molluscs from the Lower Murchison district, Southern Carnarvon Basin, Western Australia. Records of the Western Australian Museum 24 (3): 217–246.

[F27a] Finlay, H. J. 1927a. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[F27b] Finlay, H. J. 1927b. New specific names for austral Mollusca. Transactions and Proceedings of the New Zealand Institute 57: 488–533.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1–167.

[GW02] Giribet, G., & W. Wheeler. 2002. On bivalve phylogeny: a high-level analysis of the Bivalvia (Mollusca) based on combined morphology and DNA sequence data. Invertebrate Biology 121 (4): 271–324.

[G31] Grabau, A. W. 1931. The Permian of Mongolia: A report on the Permian fauna of the Jisu Honguer limestone of Mongolia and its relations to the Permian of other parts of the world. American Museum of Natural History: New York.

[H00] Hodges, P. 2000. The Early Jurassic Bivalvia from the Hettangian and Lower Sinemurian of south-west Britain. Part 1. Monograph of the Paleontographical Society 614: 1–64.

[HG98] Holte, B., & B. Gulliksen. 1998. Common macrofaunal dominant species in the sediments of some north Norwegian and Svalbard glacial fjords. Polar Biology 19: 375–382.

[JB12] Johnson, M. E., & B. G. Baarli. 2012. Development of intertidal biotas through Phanerozoic time. In: Talent, J. A. (ed.) Earth and Life: Global biodiversity, extinction intervals and biogeographic perturbations through time pp. 63–128. Springer.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O’Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

[N79] Norman, A. M. 1879. The Mollusca of the fiords near Bergen, Norway. Journal of Conchology 2: 8–77.

[N00] Norman, A. M. 1900. British Amphipoda: families Pontoporeidae to Ampeliscidae. Annals and Magazine of Natural History, series 7, 5: 326–346.

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4th ed. Whitcombe and Tombs Limited: Christchurch.

[SB93] Skelton, P. W., & M. J. Benton. 1993. Mollusca: Rostroconchia, Scaphopoda and Bivalvia. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 237–263. Chapman & Hall: London.

[XZ10] Xu, F., & J. Zhang. 2010. Study on the bivalve faunal composition of the Yellow Sea and Bohai Sea. In: China-Russia Bilateral Symposium: Proceedings of the China-Russia Bilateral Symposium of “Comparison on Marine Biodiversity in the Northwest Pacific Ocean”, 10–11 October 2010, Qingdao (China) pp. 7–12. Institute of Oceanology, Chinese Academy of Sciences; A. V. Zhirmunsky Institute of Marine Biology, Far East Branch of the Russian Academy of Sciences.

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