Paramacronychiinae

Belongs within: Oestroidea.

The Paramacronychiinae are a group of sarcophagid flies combining a non-plumose arista with strong hairs on the meron. They are diverse in habits, including necrophagous species, predators of snails and parasites of other insects.

Characters (from Pape & Dahlem 2010): Head profile with convex occipital region; antennal arista not plumose; eyes not or only slightly enlarged; postocular setae alternating in size; notopleuron without subprimary bristles; metasternal area bare; hind coxa bare posteriorly; coxopleural streak present; male midfemur with apical posteroventral ctenidium of stout, flattened bristles; male abdominal sternites 2–4 partly hidden by overlapping margins of corresponding tergites, terminalia usually protruding, with tergite 6 distinctly fused to syntergosternite 7+8 at more or less right angle.

<==ParamacronychiinaePD10
    |--Chrysogramma [Chrysogrammatini]F92
    |    `--C. variabileF92
    `--+--SarcotachiiniF92
       `--+--Goniophyto Townsend 1927 [incl. Sinagria Rohdendorf in Fan 1965; Goniophytoini]F92
          |    |--*G. formosensis Townsend 1927F92
          |    `--G. honshuensis Rohdendorf 1962F92
          `--ParamacronychiiniF92
               |--+--+--NyctiininaF92
               |  |  `--+--DexagriinaF92
               |  |     `--BlaesoxiphellinaF92
               |  `--+--Paramacronychia [Paramacronychiina]F92
               |     |    `--P. flavipalpisF92
               |     `--+--GalapagomyiinaF92
               |        `--Agria Robineau-Desvoidy 1830 [Agriina]F92
               |             |--*A. punctata Robineau-Desvoidy 1830F92
               |             |--A. houseiS87
               |             `--A. xiangchengensis Chao & Zhang 1988F92
               `--Wohlfahrtia Brauer & Bergenstamm 1889 [Wohlfahrtiina]F92
                    |--*W. magnifica (Schiner 1862) [=Sarcophila magnifica; incl. S. wohlfahrti Portschinsky 1875]F92
                    |--W. atra Aldrich 1926F92
                    |--W. balassogloi (Portschinsky 1882) [=Sarcophila balassogloi]F92
                    |--W. bella (Macquart 1838) [=Agria bella]F92
                    |--W. cheni Rohdendorf 1956F92
                    |--W. fedtschenkoi Rohdendorf 1956F92
                    |--W. intermedia (Portschinsky 1887) [=Sarcophila intermedia]F92
                    |--W. meigeni (Schiner 1862) [=Sarcophila meigeni]F92
                    |--W. nuba (Wiedemann 1830)KP10
                    |--W. pavlovskyi Rohdendorf 1956F92
                    |--W. stackelbergi Rohdendorf 1956F92
                    |--W. trina (Wiedemann 1830)KP10
                    `--W. vigilS87
Paramacronychiinae incertae sedis:
  CattasomaPD10
  Brachicoma Rondani 1856F92
    |--B. devia (Fallén 1820) (see below for synonymy)F92
    |--B. asiatica Rohdendorf & Verves 1979F92
    |--B. nigra Chao & Zhang 1988F92
    `--B. setosaPD10
  Erythandra picipesPD10, S87
  Toxonagria montanensisS87
  SarcofahrtiaS87
    |--S. montanensisE66
    `--S. raviniaS87
  Nyctia lugubris (Macquart 1843)KP10
  Sarcophila Rondani 1856KP10
    |--*S. latifrons (Fallén 1817) [=Musca latifrons]F92
    |--S. japonica (Rohdendorf 1962)I92
    |--S. meridionalis Verves 1982KP10
    |--S. mongolica Chao & Zhang 1988F92
    `--S. rasnitzyni Rohdendorf & Verves 1985F92
  Angiometopa Brauer & Bergenstamm 1889F92
    |--*A. ruralis (Fallén 1817) [=Musca ruralis]F92
    `--A. hikosanaF92
  Wohlfahrtiodes Villeneuve 1910F92
    |--*W. nudus Villeneuve 1910F92
    `--W. mongolicus Chao & Zhang 1988F92

Brachicoma devia (Fallén 1820) [=Tachina devia; incl. T. nitidula Rondani 1856 non Meigen 1824, *Brachicoma nitidula]^F92

*Type species of generic name indicated

References

[E66] Evans, H. E. 1966. The Comparative Ethology and Evolution of the Sand Wasps. Harvard University Press: Cambridge (Massachusetts).

[F92] Fan Z. 1992. Key to the Common Flies of China 2^nd ed. Science Press: Beijing.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[KP10] Kutty, S. N., T. Pape, B. M. Wiegmann & R. Meier. 2010. Molecular phylogeny of the Calyptratae (Diptera: Cyclorrhapha) with an emphasis on the superfamily Oestroidea and the position of Mystacinobiidae and McAlpine’s fly. Systematic Entomology 35: 614–635.

[PD10] Pape, T., & G. A. Dahlem. 2010. Sarcophagidae (flesh flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 1313–1335. NRC Research Press: Ottawa.

[S87] Shewell, G. E. 1987. Sarcophagidae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 1159–1186. Research Branch, Agriculture Canada.