Belongs within: Pleosporales.
The Phaeosphaeriaceae are a group of fungi that are saprobic or necrotrophic on a variety of plant species (Kirk et al. 2001).
Characters (Kirk et al. 2001): Ascomata perithecial, immersed to erumpent, gregarious or occasionally as locules in small stromata, more or less globose, with well-developed lysigenous, often periphysate ostiole; peridium soft, composed of relatively small thin-walled pseudoparenchymatous cells. Interascal tissue sparse, of narrow cellular pseudoparaphyses. Asci cylindrical, fissitunicate, with well-developed ocular chamber, not blueing in iodine. Ascospores brown, transversely septate, constricted only at primary septum, occasionally muriform, sometimes with sheath. Anamorphs coelomycetous.
<==Phaeosphaeriaceae
|--Barria Yuan 1994KC01
|--Bricookea Barr 1982KC01
|--Carinispora Hyde 1992KC01
|--Hadrospora Coise 1989KC01
|--Isthmosporella Shearer & Crane 1999KC01
|--Lautitia Schatz 1984KC01
|--Metameris Theiss. & Syd. 1915 [incl. Scirrhodothis Theiss. & Syd. 1915, Scirrhophragma Theiss. & Syd. 1915]KC01
|--Mixtura Erikss. & Yue 1990KC01
|--Nodulosphaeria Rabenh. 1858 [incl. Pocosphaeria (Sacc.) Berl. 1892]KC01
|--Paraphaeosphaeria Erikss. 1967 (see below for synonymy)KC01
|--WilmiaEB03O
|--Tiarospora Sacc. & Marchal 1885KC01
|--Eudarluca Speg. 1908 (see below for synonymy)KC01
| `--E. caricisKC01 [incl. Darluca filumKD83, Sphaerellopsis filumKC01]
|--Ophiosphaerella Speg. 1909KC01
| `--O. herpotrichaSS09
|--Phaeosphaeria Miyake 1909KC01 (see below for synonymy)
| | i. s.: ‘Sulcispora’ pleurosporaEB03N
| |--P. nodorumSS09
| `--+--P. avenariaSS09
| `--P. eustomaSS09
|--Septoria Sacc. 1884 (nom. cons.)LS01, KC01 (see below for synonymy)
| |--S. apiicolaKC01
| |--S. azaleaeKC01
| |--S. exoticaL97
| |--S. gladioliKC01
| |--S. lycopersiciKD83
| |--S. nodorumLS01
| |--S. protearumSH05
| `--S. triticiKC01
`--Tintelnotia destructansV-LC-L18
Eudarluca Speg. 1908 [incl. Botryella Syd. & Syd. 1916, Cryptocrea Petr. 1937, Darluca Castagne 1851, Darlucella Höhn. 1919, Diplodothiorella Bubák 1916, Diploplacis Clem. & Shear 1931, Diploplacosphaeria Petr. 1921, Kabathia Nieuwl. 1916, Metabotryon Syd. 1926, Mycepimyce Nieuwl. 1916, Parabotryon Syd. 1926, Sphaerellopsis Cooke 1883, Xenodimerium Petr. 1947]KC01
Paraphaeosphaeria Erikss. 1967 [incl. Chaetosphaeropsis Curzi & Barbaini 1927, Coniothyriopsis Speg. 1910, Cryptophacella Höhn. 1917, Dothisphaeropsis Höhn. 1919, Microsphaeropsis Höhn. 1917, Microsporella Höhn. 1918, Phyllostictella Tassi 1901, Sclerothyrium Höhn. 1918]KC01
Phaeosphaeria Miyake 1909KC01 [incl. Brencklea Petr. 1923KC01, Phaeoseptoria Speg. 1908KC01, Scolecosporiella Petr. 1921KC01, Sulcispora Shoemaker & Babc. 1989EB03N, KC01, Trematosphaerella Kirschst. 1906KC01]
Septoria Sacc. 1884 (nom. cons.)LS01, KC01 [=Septaria Fr. 1821 (nom. rej.) non Férussac 1807 (ICZN)KC01; incl. Septoriopsis Gonz. Frag. & Paúl 1915KC01, Spilosphaeria Rabenh. 1857KC01]
*Type species of generic name indicated
References
[EB03N] Eriksson, O. E., H. O. Barah, R. S. Currah, K. Hansen, C. P. Kurtzman, T. Laessøe & G. Rambold (eds.) 2003. Notes on ascomycete systematics. Nos 3580–3623. Myconet 9: 91–103.
[EB03O] Eriksson, O. E., H. O. Barah, R. S. Currah, K. Hansen, C. P. Kurtzman, G. Rambold & T. Laessøe (eds.) 2003. Outline of Ascomycota—2003. Myconet 9: 1–89.
[KC01] Kirk, P. M., P. F. Cannon, J. C. David & J. A. Stalpers. 2001. Ainsworth & Bisby’s Dictionary of the Fungi 9th ed. CAB International: Wallingford (UK).
[KD83] Koç, N. K., & G. Défago. 1983. Studies on the host range of the hyperparasite Aphanocladium album. Phytopathologische Zeitschrift 107: 214–218.
[L97] Lange, P. J. de. 1997. Hebe brevifolia (Scrophulariaceae)—an ultramafic endemic of the Surville Cliffs, North Cape, New Zealand. New Zealand Journal of Botany 35: 1–8.
[LS01] Lumbsch, H. T., I. Schmitt, H. Döring & M. Wedin. 2001. Molecular systematics supports the recognition of an additional order of Ascomycota: the Agyriales. Mycological Research 105 (1): 16–23.
[SS09] Schoch, C. L., G.-H. Sung, F. López-Giráldez, J. P. Townsend, J. Miadlikowska, V. Hofstetter, B. Robbertse, P. B. Matheny, F. Kauff, Z. Wang, C. Gueidan, R. M. Andrie, K. Trippe, L. M. Ciufetti, A. Wynns, E. Fraker, B. P. Hodkinson, G. Bonito, J. Z. Groenewald, M. Arzanlou, G. S. de Hoog, P. W. Crous, D. Hewitt, D. H. Pfister, K. Peterson, M. Gryzenhout, M. J. Wingfield, A. Aptroot, S.-O. Suh, M. Blackwell, D. M. Hillis, G. W. Griffith, L. A. Castlebury, A. Y. Rossman, H. T. Lumbsch, R. Lücking, B. Büdel, A. Rauhut, P. Diederich, D. Ertz, D. M. Geiser, K. Hosaka, P. Inderbitzin, J. Kohlmeyer, B. Volkmann-Kohlmeyer, L. Mostert, K. O’Donnell, H. Sipman, J. D. Rogers, R. A. Shoemaker, J. Sugiyama, R. C. Summerbell, W. Untereiner, P. R. Johnston, S. Stenroos, A. Zuccaro, P. S. Dyer, P. D. Crittenden, M. S. Cole, K. Hansen, J. M. Trappe, R. Yahr, F. Lutzoni & J. W. Spatafora. 2009. The Ascomycota tree of life: a phylum-wide phylogeny clarifies the origin and evolution of fundamental reproductive and ecological traits. Systematic Biology 58 (2): 224–239.
[SH05] Selbmann, L., G. S. de Hoog, A. Mazzaglia, E. I. Friedman & S. Onofri. 2005. Fungi at the edge of life: cryptoendolithic black fungi from Antarctic desert. Studies in Mycology 51: 1–32.
[V-LC-L18] Valenzuela-Lopez, N., J. F. Cano-Lira, A. M. Stchigel & J. Guarro. 2018. DNA sequencing to clarify the taxonomical conundrum of the clinical coelomycetes. Mycoses 61: 708–717.