Pholidotomoidea

Napulus fragilis, from fossilshells.nl.

Belongs within: Neogastropoda.
Contains: Pyrifusinae, Volutodermatinae, Beretrinae.

The Pholidotomoidea as recognised by Bouchet et al. (2017) include early representatives of the Neogastropoda from the Late Cretaceous. This group is probably not monophyletic and likely includes ancestors of later neogastropods.

<==Pholidotomoidea [Pyrifusoidea, Sarganoidea]BR17
|--WeeksiidaeBD12
| |--‘Lowenstamia’ Sohl 1964 non Stehli 1961BD12
| | `--*L. funiculus Sohl 1964RA75
| `--Weeksia Stephenson 1941BR05
| |--*W. amplificata (Wade 1926) [=Pseudomalaxis amplificata]KC60
| |--W. deplanata (Johnson 1905)TTE93
| `--W. lubbocki (Stephenson 1941)TTE93
|--Pholidotomidae [Pyrifusidae]BR17
| |--PyrifusinaeBD12
| |--VolutodermatinaeBD12
| |--BeretrinaeBR17
| |--Paleopsephaea Wade 1926BD12 [PaleopsephaeinaeBR17]
| | `--*P. mutabilis Wade 1926BD12
| |--Pseudorapa Holzapfel 1888 [Pseudorapinae]BD12
| | |--*P. (Pseudorapa) pleurotomoides (Muller 1851) [=Murex pleurotomoides]RA75
| | |--P. (*Aliofusus) stamineus Sohl 1964BD12
| | `--P. (Stantonella Wade 1926)BD12
| | |--P. (*S.) subnodosa (Wade 1926) [=*Stantonella subnodosa]BD12
| | `--P. (S.) ripleyana (Conrad 1860)BD12
| `--PholidotominaeBD12
| |--Pholidotoma Cossmann 1896BD12
| | `--*P. subheptagonus (d’Orbigny 1850) [=Fusus subheptagonus]BR17
| `--Drilluta Wade 1916BD12 [=Drilliovoluta Cossmann 1925P66; Drillutinae]
| |--*D. communis Wade 1916BD12 [=*Drilliovoluta communisP66]
| |--D. distans (Conrad 1860)BD12
| `--D. majorBD12
|--SarganidaeBR17
| |--Hippocampoides Wade 1916RA75 [HippocampoidinaeBD12]
| | `--*H. serratus Wade 1916RA75
| |--SchizobasinaeBD12
| | |--Hillites Stephenson 1952BD12
| | |--‘Natalites’ Bandel 1999 non Collignon 1954BD12
| | `--Schizobasis Wade 1916 non Baker 1873 (ICBN)BR05
| | `--*S. depressa Wade 1916BR17
| |--SarganinaeBD12
| | |--Sargana Stephenson 1923RA75
| | | `--*S. stantoni (Stephenson 1923) [=Rapana stantoni]RA75
| | `--Praesargana Saul & Popenoe 1993BD12
| | |--*P. condoni (White 1889) [=Trophon condoni]BD12
| | `--P. tupeloensis Bandel & Dockery 2001BD12
| `--PseudecphorinaeBD12
| | i. s.: ‘Trigonostoma’ ripleyana Sohl 1964BD12
| |--Pseudecphora Bandel & Dockery 2001BR17
| | `--*P. proquadricostata (Wade 1917) [=Ecphora proquadricostata]BR17
| `--Blackdownia Kollmann 1976BD12
| |--*B. quadrata (Sowerby 1823) [=Murex quadratus]BD12
| |--B. acuticarinata (Rennie 1930)BD12
| `--B. kollmanni Kiel & Bandel 2003BD12
`--MoreidaeBD12
|--MoreinaeBD12
| |--Paramorea Wade 1917BD12
| | |--P. lineata (Sowerby 1836)TTE93
| | `--P. lirata Wade 1917BD12
| |--Morea Conrad 1860BR05 [incl. Pseudomorea Cossmann 1925RA75]
| | |--*M. cancellaria Conrad 1860RA75
| | `--M. marylandica Gardner 1916BD12
| `--Buccinopsis Conrad 1857 [incl. Seminola Wade 1918]BD12
| |--*B. perryi Conrad 1857BD12
| `--B. solida (Wade 1917)BD12
`--Pyropsinae [Pyropsidae]BD12
|--Hydrotribulus Wade 1916BD12
|--Trochifusus Wade 1926BS00
| |--*T. perornatus Wade 1926 [=Pyropsis perornatus]BS00
| |--T. africanus (Woods 1906)BS00
| `--T. hombronianus (Orbigny 1846) [=Pyrula hombroniana, Tudicla (Pyropsis) hombroniana]BS00
|--Hercorhyncus Conrad 1868BD12
| `--*H. tippanus (Conrad 1860) [=Fusus tippana]BD12
|--Schizofusus Kiel & Bandel 2003BD12
| `--*S. transkeiensis Kiel & Bandel 2003BD12
|--Lupira Stephenson 1941BD12
| `--*L. variabilis (Wade 1926) [=Xancus variabilis]BD12
|--Pyropsis Conrad 1860BR05
| | i. s.: P. bairdii Meek & Hayden 1857BS00
| | P. nodifera (Stoliczka 1868)BD12
| | ‘Tudicla’ rugosa (Wetzel 1930) non Pyrula rugosa Philippi 1887BS00
| |--P. (Pyropsis)BS00
| | |--*P. (P.) perlata (Conrad 1860) [=Tudicla perlata]BR17
| | `--P. (P.) rugosa (Philippi 1887) [=Pyrula rugosa]BS00
| `--P. (Chilenopsis Bandel & Stinnesbeck 2000)BS00
| `--P. (*C.) quinzioensis Bandel & Stinnesbeck 2000BS00
`--Napulus Stephenson 1941BD12
|--N. excavatus Stoliczka 1868BD12
|--N. fragilisBD12
|--N. reesidei Sohl 1964BD12
`--N. tuberculatus Stephenson 1941BD12

*Type species of generic name indicated

References

[BD12] Bandel, K., & D. T. Dockery III. 2012. Protoconch characters of Late Cretaceous Latrogastropoda (Neogastropoda and Neomesogastropoda) as an aid in the reconstruction of the phylogeny of the Neogastropoda. Freiberger Forschungshefte C542 (psf 20): 93–128.

[BS00] Bandel, K., & W. Stinnesbeck. 2000. Gastropods of the Quiriquina Formation (Maastrichtian) in central Chile: paleobiogeographic relationships and the description of a few new taxa. Zbl. Geol. Paläont. Teil I 7–8: 757–788.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I169–I331. Geological Society of America, and University of Kansas Press.

[P66] Powell, A. W. B. 1966. The molluscan families Speightiidae and Turridae: an evaluation of the valid taxa, both Recent and fossil, with lists of characteristic species. Bulletin of the Auckland Institute and Museum 5: 1–184, pls 1–23.

[RA75] Radwin, G. E., & A. D’Attilio. 1975. A catalogue of muricacean generic taxa. Transactions of the San Diego Society of Natural History 17 (20): 279–292.

[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.

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