Varuninae

Belongs within: Grapsoidea.

The Varuninae are a group of coastal crabs with a subquadrate or oval carapace.

<==Varuninae [Varunidae]
    |--Helograpsus haswellianus (Whitelegge 1899)SCF02
    |--Metaplax elegans de Man 1888SCF02
    |--Paragrapsus laevis (Dana 1851)SCF02
    |--MiersiograpsusSCF02
    |--XenograpsusSCF02
    |--Pseudograpsus elongatus (Milne Edwards 1873)HS15
    |--PtychognathusCC03
    |--Neohelice granulata (Dana 1851) [=Chasmagnathus granulatus]GN-F08
    |--Varuna Milne Edwards 1830BWW93
    |    `--V. litterata (Fabricius 1798)CC03
    |--Helice de Haan 1835SCF02, B64
    |    `--H. crassa Dana 1851SCF02 [incl. H. lucasi Milne Edwards 1853B64]
    |--Heterograpsus Lucas 1846B64, E12
    |    `--H. sexdentatus Lucas 1846E12
    |--CyrtograpsusSCF02
    |    |--C. affinis (Dana 1851)SCF02
    |    `--C. angulatus Dana 1851GN-F08
    |--Brachynotus de Haan 1835 [incl. Leptograpsodes Montgomery 1931]B64
    |    |--B. atlanticus Forest 1957SCF02
    |    |--B. foresti Zariquiey-Alvarez 1968KK03
    |    |--B. gemellari (Rizza 1839)KK03
    |    |--B. octodentatus [incl. Leptograpsodes webhaysi]B64
    |    `--B. sexdentatus (Risso 1826) [=Goneplax sexdentatus]B64
    |--Eriocheir De Haan 1835 [incl. Neoeriocheir Sakai 1983, Platyeriocheir Ng et al. 1999]CC03
    |    |--E. leptognathus Rathbun 1913 [=Neoeriocheir leptognathus]CC03
    |    `--+--E. formosa Chan et al. 1995 [=Platyeriocheir formosa]CC03
    |       `--+--*E. japonica (De Haan 1835) [incl. E. recta (Stimpson 1858)]CC03
    |          |--E. hepuensis Dai 1991 [=E. japonica hepuensis]CC03
    |          `--E. sinensis Milne Edwards 1854CC03
    |--Hemigrapsus Dana 1851B64
    |    |--H. crenulatus (Milne Edwards 1837) [=Cyclograpsus crenulatus, Brachynotus crenulatus]B64
    |    |--H. granarius (Nicolet 1849) [=Trichodactylus granarius; incl. H. granulatus]B64
    |    |--H. maculatusB64
    |    |--H. oregonensis (Dana 1851)SCF02
    |    |--H. penicillatusGLT03
    |    |--H. sanguineusGLT03
    |    `--H. sexdentatus (Milne Edwards 1837) (see below for synonymy)B64
    `--OrcovitaJG19
         |--O. hicksi Davie & Ng 2012JG19
         `--O. orchardorum Davie & Ng 2012JG19

Hemigrapsus sexdentatus (Milne Edwards 1837) [=Cyclograpsus sexdentatus; incl. Brachynotus edwardsi Hilgendorf 1882, Hemigrapsus edwardsi]^B64

*Type species of generic name indicated

References

[B64] Bennett, E. W. 1964. The marine fauna of New Zealand: Crustacea Brachyura. New Zealand Department of Scientific and Industrial Research Bulletin 153: 1–120.

[BWW93] Briggs, D. E. G., M. J. Weedon & M. A. Whyte. 1993. Arthropoda (Crustacea excluding Ostracoda). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 321–342. Chapman & Hall: London.

[CC03] Chu, K. H., H. Y. Cho, C. P. Li & T.-Y. Chan. 2003. Molecular phylogenetics of the mitten crab species in Eriocheir, sensu lato (Brachyura: Grapsidae). Journal of Crustacean Biology 23 (3): 738–746.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[GLT03] Glenner, H., J. Lützen & T. Takahashi. 2003. Molecular and morphological evidence for a monophyletic clade of asexually reproducing Rhizocephala: Polyascus, new genus (Cirripedia). Journal of Crustacean Biology 23: 548–557.

[GN-F08] Gregati, R. A., & M. L. Negreiros-Franzoso. 2008. Distribution of Neohelice granulata (Dana 1851) in its northernmost latitudinal occurrence in a tropical mangrove forest, Brazil (Crustacea, Decapoda, Brachyura, Grapsoidea, Varunidae). Senckenbergiana Biologica 88 (2): 177–184.

[HS15] Hosie, A. M., A. Sampey, P. J. F. Davie & D. S. Jones. 2015. Kimberley marine biota. Historical data: crustaceans. Records of the Western Australian Museum Supplement 84: 247–285.

[JG19] James, D. J., P. T. Green, W. F. Humphreys & J. C. Z. Woinarski. 2019. Endemic species of Christmas Island, Indian Ocean. Records of the Western Australian Museum 34 (2): 55–114.

[KK03] Kocataş, A., & T. Katağan. 2003. The decapod crustacean fauna of the Turkish seas. Zoology in the Middle East 29: 63–74.

[SCF02] Schubart, C. D., J. A. Cuesta & D. L. Felder. 2002. Glyptograpsidae, a new brachyuran family from Central America: larval and adult morphology, and a molecular phylogeny of the Grapsoidea. Journal of Crustacean Biology 22 (1): 28–44.